{
  "pipeline_id": "tree-builder-native-vs-iqtree2-54taxa-calibration-v1",
  "pipeline_title": "54-Taxon (106-Node) Jawed Vertebrate Phylogeny Calibration: Native ML Tree Builder (BIONJ+NNI) vs. IQ-TREE v2.4.0 Across 8 Substitution & Search Models (Using Native ModelTest Only)",
  "data_provenance": "EMPIRICAL DATA: NCBI REFSEQ/GENBANK CACHED (54 COMPLETE VERTEBRATE MITOGENOMES)",
  "modeltest_policy": "Native ModelTest Only (ModelTestMode::Native — Column-Compressed Felsenstein Pruning)",
  "total_duration_ms": 273200,
  "phylogeny_title": "Gnathostomata (Jawed Vertebrates: Chondrichthyes, Actinopterygii, Amphibia, Reptilia, Aves, Mammalia) Mitochondrial COX1 (1,545 bp)",
  "leaf_taxa_count": 55,
  "internal_node_count": 53,
  "total_tree_nodes": 108,
  "nontrivial_bipartitions": 52,
  "alignment_length_bp": 1545,
  "specimen_ledger": [
    {
      "tip_label": "Carcharodon_carcharias",
      "accession": "NC_022415.1",
      "species": "Carcharodon carcharias",
      "common_name": "Great white shark",
      "vertebrate_class": "Chondrichthyes",
      "clade_order": "Lamniformes"
    },
    {
      "tip_label": "Isurus_oxyrinchus",
      "accession": "NC_022691.1",
      "species": "Isurus oxyrinchus",
      "common_name": "Shortfin mako shark",
      "vertebrate_class": "Chondrichthyes",
      "clade_order": "Lamniformes"
    },
    {
      "tip_label": "Isurus_paucus",
      "accession": "NC_024101.1",
      "species": "Isurus paucus",
      "common_name": "Longfin mako shark",
      "vertebrate_class": "Chondrichthyes",
      "clade_order": "Lamniformes"
    },
    {
      "tip_label": "Carcharhinus_acronotus",
      "accession": "NC_024055.1",
      "species": "Carcharhinus acronotus",
      "common_name": "Blacknose shark",
      "vertebrate_class": "Chondrichthyes",
      "clade_order": "Carcharhiniformes"
    },
    {
      "tip_label": "Okamejei_kenojei",
      "accession": "NC_007173.1",
      "species": "Okamejei kenojei",
      "common_name": "Ocellate spot skate",
      "vertebrate_class": "Chondrichthyes",
      "clade_order": "Rajiformes"
    },
    {
      "tip_label": "Danio_rerio",
      "accession": "NC_002333.2",
      "species": "Danio rerio",
      "common_name": "Zebrafish",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Cypriniformes"
    },
    {
      "tip_label": "Cyprinus_carpio",
      "accession": "NC_001606.1",
      "species": "Cyprinus carpio",
      "common_name": "Common carp",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Cypriniformes"
    },
    {
      "tip_label": "Salvelinus_fontinalis",
      "accession": "NC_000860.1",
      "species": "Salvelinus fontinalis",
      "common_name": "Brook trout",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Salmoniformes"
    },
    {
      "tip_label": "Oncorhynchus_mykiss",
      "accession": "NC_001717.1",
      "species": "Oncorhynchus mykiss",
      "common_name": "Rainbow trout",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Salmoniformes"
    },
    {
      "tip_label": "Gadus_morhua",
      "accession": "NC_002081.1",
      "species": "Gadus morhua",
      "common_name": "Atlantic cod",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Gadiformes"
    },
    {
      "tip_label": "Plectorhinchus_lineatus",
      "accession": "NC_025517.1",
      "species": "Plectorhinchus lineatus",
      "common_name": "Lined sweetlips",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Perciformes"
    },
    {
      "tip_label": "Thunnus_thynnus",
      "accession": "NC_014052.1",
      "species": "Thunnus thynnus",
      "common_name": "Atlantic bluefin tuna",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Scombriformes"
    },
    {
      "tip_label": "Thunnus_orientalis",
      "accession": "NC_008455.1",
      "species": "Thunnus orientalis",
      "common_name": "Pacific bluefin tuna",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Scombriformes"
    },
    {
      "tip_label": "Thunnus_albacares",
      "accession": "NC_014061.1",
      "species": "Thunnus albacares",
      "common_name": "Yellowfin tuna",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Scombriformes"
    },
    {
      "tip_label": "Thunnus_maccoyii",
      "accession": "NC_014101.1",
      "species": "Thunnus maccoyii",
      "common_name": "Southern bluefin tuna",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Scombriformes"
    },
    {
      "tip_label": "Thunnus_alalunga",
      "accession": "NC_005317.1",
      "species": "Thunnus alalunga",
      "common_name": "Albacore tuna",
      "vertebrate_class": "Actinopterygii",
      "clade_order": "Scombriformes"
    },
    {
      "tip_label": "Xenopus_laevis",
      "accession": "NC_001573.1",
      "species": "Xenopus laevis",
      "common_name": "African clawed frog",
      "vertebrate_class": "Amphibia",
      "clade_order": "Anura"
    },
    {
      "tip_label": "Chelonia_mydas",
      "accession": "NC_000886.1",
      "species": "Chelonia mydas",
      "common_name": "Green sea turtle",
      "vertebrate_class": "Reptilia",
      "clade_order": "Testudines"
    },
    {
      "tip_label": "Alligator_mississippiensis",
      "accession": "NC_001922.1",
      "species": "Alligator mississippiensis",
      "common_name": "American alligator",
      "vertebrate_class": "Reptilia",
      "clade_order": "Crocodylia"
    },
    {
      "tip_label": "Struthio_camelus",
      "accession": "NC_002785.1",
      "species": "Struthio camelus",
      "common_name": "Common ostrich",
      "vertebrate_class": "Aves",
      "clade_order": "Struthioniformes"
    },
    {
      "tip_label": "Gallus_gallus",
      "accession": "AP003322.1",
      "species": "Gallus gallus",
      "common_name": "Red junglefowl",
      "vertebrate_class": "Aves",
      "clade_order": "Galliformes"
    },
    {
      "tip_label": "Meleagris_gallopavo",
      "accession": "NC_010195.2",
      "species": "Meleagris gallopavo",
      "common_name": "Wild turkey",
      "vertebrate_class": "Aves",
      "clade_order": "Galliformes"
    },
    {
      "tip_label": "Coturnix_japonica",
      "accession": "NC_003408.1",
      "species": "Coturnix japonica",
      "common_name": "Japanese quail",
      "vertebrate_class": "Aves",
      "clade_order": "Galliformes"
    },
    {
      "tip_label": "Anas_platyrhynchos",
      "accession": "NC_009684.1",
      "species": "Anas platyrhynchos",
      "common_name": "Mallard duck",
      "vertebrate_class": "Aves",
      "clade_order": "Anseriformes"
    },
    {
      "tip_label": "Columba_livia",
      "accession": "NC_013978.1",
      "species": "Columba livia",
      "common_name": "Rock dove",
      "vertebrate_class": "Aves",
      "clade_order": "Columbiformes"
    },
    {
      "tip_label": "Falco_peregrinus",
      "accession": "NC_000878.1",
      "species": "Falco peregrinus",
      "common_name": "Peregrine falcon",
      "vertebrate_class": "Aves",
      "clade_order": "Falconiformes"
    },
    {
      "tip_label": "Ornithorhynchus_anatinus",
      "accession": "NC_000891.1",
      "species": "Ornithorhynchus anatinus",
      "common_name": "Platypus",
      "vertebrate_class": "Mammalia",
      "clade_order": "Monotremata"
    },
    {
      "tip_label": "Monodelphis_domestica",
      "accession": "NC_006299.1",
      "species": "Monodelphis domestica",
      "common_name": "Gray short-tailed opossum",
      "vertebrate_class": "Mammalia",
      "clade_order": "Didelphimorphia"
    },
    {
      "tip_label": "Loxodonta_africana",
      "accession": "NC_000934.1",
      "species": "Loxodonta africana",
      "common_name": "African savanna elephant",
      "vertebrate_class": "Mammalia",
      "clade_order": "Proboscidea"
    },
    {
      "tip_label": "Oryctolagus_cuniculus",
      "accession": "NC_001913.1",
      "species": "Oryctolagus cuniculus",
      "common_name": "European rabbit",
      "vertebrate_class": "Mammalia",
      "clade_order": "Lagomorpha"
    },
    {
      "tip_label": "Mus_musculus",
      "accession": "NC_005089.1",
      "species": "Mus musculus",
      "common_name": "House mouse",
      "vertebrate_class": "Mammalia",
      "clade_order": "Rodentia"
    },
    {
      "tip_label": "Rattus_norvegicus",
      "accession": "NC_001665.2",
      "species": "Rattus norvegicus",
      "common_name": "Brown rat",
      "vertebrate_class": "Mammalia",
      "clade_order": "Rodentia"
    },
    {
      "tip_label": "Macaca_mulatta",
      "accession": "NC_005943.1",
      "species": "Macaca mulatta",
      "common_name": "Rhesus macaque",
      "vertebrate_class": "Mammalia",
      "clade_order": "Primates"
    },
    {
      "tip_label": "Pongo_pygmaeus",
      "accession": "NC_001646.1",
      "species": "Pongo pygmaeus",
      "common_name": "Bornean orangutan",
      "vertebrate_class": "Mammalia",
      "clade_order": "Primates"
    },
    {
      "tip_label": "Gorilla_gorilla",
      "accession": "NC_001645.1",
      "species": "Gorilla gorilla",
      "common_name": "Western gorilla",
      "vertebrate_class": "Mammalia",
      "clade_order": "Primates"
    },
    {
      "tip_label": "Pan_troglodytes",
      "accession": "NC_001643.1",
      "species": "Pan troglodytes",
      "common_name": "Chimpanzee",
      "vertebrate_class": "Mammalia",
      "clade_order": "Primates"
    },
    {
      "tip_label": "Homo_sapiens",
      "accession": "NC_012920.1",
      "species": "Homo sapiens",
      "common_name": "Human (rCRS RefSeq)",
      "vertebrate_class": "Mammalia",
      "clade_order": "Primates"
    },
    {
      "tip_label": "Bos_taurus",
      "accession": "NC_006853.1",
      "species": "Bos taurus",
      "common_name": "Cattle",
      "vertebrate_class": "Mammalia",
      "clade_order": "Cetartiodactyla"
    },
    {
      "tip_label": "Ovis_aries",
      "accession": "NC_001941.1",
      "species": "Ovis aries",
      "common_name": "Domestic sheep",
      "vertebrate_class": "Mammalia",
      "clade_order": "Cetartiodactyla"
    },
    {
      "tip_label": "Sus_scrofa",
      "accession": "NC_000845.1",
      "species": "Sus scrofa",
      "common_name": "Wild boar",
      "vertebrate_class": "Mammalia",
      "clade_order": "Cetartiodactyla"
    },
    {
      "tip_label": "Balaenoptera_musculus",
      "accession": "NC_001601.1",
      "species": "Balaenoptera musculus",
      "common_name": "Blue whale",
      "vertebrate_class": "Mammalia",
      "clade_order": "Cetartiodactyla"
    },
    {
      "tip_label": "Equus_caballus",
      "accession": "NC_001640.1",
      "species": "Equus caballus",
      "common_name": "Domestic horse",
      "vertebrate_class": "Mammalia",
      "clade_order": "Perissodactyla"
    },
    {
      "tip_label": "Felis_catus",
      "accession": "NC_001700.1",
      "species": "Felis catus",
      "common_name": "Domestic cat",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Felidae)"
    },
    {
      "tip_label": "Panthera_pardus",
      "accession": "NC_010641.1",
      "species": "Panthera pardus",
      "common_name": "Leopard",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Felidae)"
    },
    {
      "tip_label": "Panthera_tigris_refseq",
      "accession": "NC_010642.1",
      "species": "Panthera tigris",
      "common_name": "Tiger (RefSeq)",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Felidae)"
    },
    {
      "tip_label": "Panthera_tigris_amur",
      "accession": "EF551003.1",
      "species": "Panthera tigris altaica",
      "common_name": "Amur tiger",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Felidae)"
    },
    {
      "tip_label": "Canis_lupus",
      "accession": "NC_008092.1",
      "species": "Canis lupus",
      "common_name": "Gray wolf",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Canidae)"
    },
    {
      "tip_label": "Canis_lupus_familiaris",
      "accession": "NC_002008.4",
      "species": "Canis lupus familiaris",
      "common_name": "Domestic dog",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Canidae)"
    },
    {
      "tip_label": "Vulpes_vulpes",
      "accession": "NC_008434.1",
      "species": "Vulpes vulpes",
      "common_name": "Red fox",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Canidae)"
    },
    {
      "tip_label": "Meles_anakuma",
      "accession": "NC_009677.1",
      "species": "Meles anakuma",
      "common_name": "Japanese badger",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Mustelidae)"
    },
    {
      "tip_label": "Mustela_putorius",
      "accession": "NC_020638.1",
      "species": "Mustela putorius",
      "common_name": "European polecat",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Mustelidae)"
    },
    {
      "tip_label": "Ursus_thibetanus",
      "accession": "NC_009971.1",
      "species": "Ursus thibetanus",
      "common_name": "Asian black bear",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Ursidae)"
    },
    {
      "tip_label": "Ursus_americanus",
      "accession": "NC_003426.1",
      "species": "Ursus americanus",
      "common_name": "American black bear",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Ursidae)"
    },
    {
      "tip_label": "Ursus_arctos",
      "accession": "NC_003427.1",
      "species": "Ursus arctos",
      "common_name": "Brown bear",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Ursidae)"
    },
    {
      "tip_label": "Ursus_maritimus",
      "accession": "NC_003428.1",
      "species": "Ursus maritimus",
      "common_name": "Polar bear",
      "vertebrate_class": "Mammalia",
      "clade_order": "Carnivora (Ursidae)"
    }
  ],
  "native_modeltest_prescreen": {
    "engine_used": "Native Rust Felsenstein ModelTest (ModelTestMode::Native ONLY)",
    "duration_ms": 1138.9989,
    "alignment_length_bp": 1545,
    "unique_site_patterns": 779,
    "invariant_site_fraction": 0.488,
    "best_model_bic": "HKY+I+G",
    "best_model_aic": "HKY+I+G",
    "best_bic_ln_l": -32554.7296,
    "best_bic_score": 65939.1932,
    "empirical_pi": [
      0.2691,
      0.2619,
      0.1723,
      0.2967
    ],
    "kappa_pyrimidine_ct": 3.093,
    "kappa_purine_ag": 1.3449,
    "overall_ti_tv_kappa": 2.219,
    "pinv_estimate": 0.4002,
    "gamma_alpha_estimate": 0.42
  },
  "run_model_comparisons": [
    {
      "run_model_code": "JC",
      "run_model_description": "Jukes-Cantor (1969): Equal base frequencies (0.25), single substitution rate, uniform site rates",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI)",
        "run_model_code": "JC",
        "duration_ms": 614.8477,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -40809.0422,
        "neg_ln_l": 40809.0422,
        "free_parameters_k": 105,
        "aic": 81828.0844,
        "bic": 82389.0762,
        "total_tree_length": 5.8975,
        "mean_internal_branch_length": 0.0356,
        "nni_moves_evaluated": 814,
        "nni_moves_accepted": 51,
        "mean_branch_support_pct": 99.8006,
        "newick_tree": "(((((((Oryctolagus_cuniculus:0.11687,(Monodelphis_domestica:0.12299,Ornithorhynchus_anatinus:0.14804)99:0.03209)100:0.02820,((((Okamejei_kenojei:0.14135,(Carcharhinus_acronotus:0.10060,(Carcharodon_carcharias:0.06147,(Isurus_oxyrinchus:0.05960,Isurus_paucus:0.06467)100:0.03428)100:0.05084)100:0.03885)100:0.03881,((Danio_rerio:0.09799,Cyprinus_carpio:0.09875)100:0.04578,((Plectorhinchus_lineatus:0.13049,((Thunnus_albacares:0.00000,Thunnus_maccoyii:0.00000)96:0.00087,(Thunnus_thynnus:0.00476,(Thunnus_orientalis:0.00239,Thunnus_alalunga:0.00084)100:0.01049)100:0.00348)100:0.07875)100:0.03102,(Gadus_morhua:0.11684,(Salvelinus_fontinalis:0.05752,Oncorhynchus_mykiss:0.05834)100:0.07478)100:0.03262)100:0.02094)100:0.03712)100:0.02612,(Chelonia_mydas:0.11851,(Alligator_mississippiensis:0.14386,(((Anas_platyrhynchos:0.10798,Columba_livia:0.07648)100:0.02542,(Falco_peregrinus:0.07951,Struthio_camelus:0.11274)99:0.01842)100:0.02267,(Coturnix_japonica:0.06869,(Gallus_gallus:0.06309,Meleagris_gallopavo:0.08108)100:0.02425)100:0.04048)100:0.05220)99:0.02404)100:0.03356)100:0.02508,Xenopus_laevis:0.16913)100:0.04008)100:0.02586,((Mus_musculus:0.09426,Rattus_norvegicus:0.09366)100:0.05277,((Meles_anakuma:0.08946,Mustela_putorius:0.11071)100:0.02965,((Felis_catus:0.06553,((Panthera_tigris_refseq:0.00000,Panthera_tigris_amur:0.00000)100:0.06081,Panthera_pardus:0.04315)100:0.02712)100:0.07339,(((Ursus_arctos:0.00929,Ursus_maritimus:0.00773)100:0.03698,(Ursus_thibetanus:0.04040,Ursus_americanus:0.03636)100:0.01600)100:0.07564,(Vulpes_vulpes:0.09429,(Canis_lupus:0.00098,Canis_lupus_familiaris:0.00226)100:0.07470)100:0.05309)100:0.03189)100:0.02153)100:0.02054)100:0.02443)100:0.01712,(Balaenoptera_musculus:0.12137,(Bos_taurus:0.08733,Ovis_aries:0.06287)100:0.03501)100:0.02985)100:0.01841,(Sus_scrofa:0.09497,Loxodonta_africana:0.15209)100:0.03590)100:0.02830,(Macaca_mulatta:0.10341,(Pongo_pygmaeus:0.08258,(Homo_sapiens:0.04381,(Pan_troglodytes:0.03945,Gorilla_gorilla:0.06698)97:0.01668)100:0.04027)100:0.05084)100:0.05748)100:0.03879,Equus_caballus:0.06563);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "JC",
        "duration_ms": 98.7122,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -41418.2749,
        "neg_ln_l": 41418.2749,
        "free_parameters_k": 105,
        "aic": 83046.5499,
        "bic": 83607.5417,
        "total_tree_length": 6.1244,
        "mean_internal_branch_length": 0.0217,
        "nni_moves_evaluated": 210,
        "nni_moves_accepted": 7,
        "mean_branch_support_pct": 99.1806,
        "newick_tree": "((((Chelonia_mydas:0.14531,(((Coturnix_japonica:0.09063,(Gallus_gallus:0.09061,Meleagris_gallopavo:0.09011)100:0.00574)100:0.03163,(Struthio_camelus:0.12329,(Falco_peregrinus:0.10796,(Anas_platyrhynchos:0.12156,Columba_livia:0.10793)100:0.00926)100:0.00324)100:0.00232)100:0.03055,Alligator_mississippiensis:0.17953)92:0.00113)100:0.01507,(Xenopus_laevis:0.18712,((Okamejei_kenojei:0.16611,(Carcharhinus_acronotus:0.12469,(Carcharodon_carcharias:0.08961,(Isurus_oxyrinchus:0.07790,Isurus_paucus:0.07837)100:0.01992)100:0.04596)100:0.00992)100:0.02256,((Danio_rerio:0.11921,Cyprinus_carpio:0.12183)100:0.02250,((Salvelinus_fontinalis:0.07283,Oncorhynchus_mykiss:0.07002)100:0.07517,(Gadus_morhua:0.14002,(Plectorhinchus_lineatus:0.13459,(((Thunnus_thynnus:0.00482,(Thunnus_orientalis:0.00274,Thunnus_alalunga:0.00136)100:0.00875)95:0.00363,Thunnus_maccoyii:0.00025)96:0.00025,Thunnus_albacares:0.00253)100:0.11671)99:0.00672)100:0.00374)100:0.00630)100:0.01744)100:0.00902)99:0.00422)100:0.02404,(Monodelphis_domestica:0.15848,(Ornithorhynchus_anatinus:0.17038,Loxodonta_africana:0.16406)95:0.00537)96:0.00204)93:0.00058,(Oryctolagus_cuniculus:0.14446,((Balaenoptera_musculus:0.13806,(Bos_taurus:0.09768,Ovis_aries:0.09000)100:0.02247)100:0.00950,(((Meles_anakuma:0.11872,((Mus_musculus:0.11442,Rattus_norvegicus:0.11716)100:0.02602,Mustela_putorius:0.13186)100:0.01276)100:0.00752,((Felis_catus:0.08615,(Panthera_pardus:0.06593,(Panthera_tigris_refseq:0.00025,Panthera_tigris_amur:0.00025)100:0.06720)100:0.01944)100:0.06007,(((Ursus_arctos:0.01289,Ursus_maritimus:0.00863)100:0.04942,(Ursus_thibetanus:0.05011,Ursus_americanus:0.04813)100:0.00995)100:0.07697,(Vulpes_vulpes:0.10444,(Canis_lupus:0.00135,Canis_lupus_familiaris:0.00275)100:0.10465)100:0.02978)100:0.00854)100:0.00578)100:0.00440,(Sus_scrofa:0.13014,(Equus_caballus:0.12982,(Macaca_mulatta:0.13510,(Pongo_pygmaeus:0.11074,(Gorilla_gorilla:0.07259,(Pan_troglodytes:0.05764,Homo_sapiens:0.05893)97:0.01147)100:0.02629)100:0.03992)100:0.03422)99:0.00644)100:0.00667)100:0.00364)100:0.00486)96:0.00058);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "JC",
        "duration_ms": 15013.6446,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -40787.8694,
        "neg_ln_l": 40787.8694,
        "free_parameters_k": 107,
        "aic": 81789.7389,
        "bic": 82361.4162,
        "total_tree_length": 5.8794,
        "mean_internal_branch_length": 0.0342,
        "nni_moves_evaluated": 954,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 82.0231,
        "newick_tree": "(Carcharodon_carcharias:0.0615637089,(Isurus_oxyrinchus:0.0597324321,Isurus_paucus:0.0644305325)99.7:0.0344629969,(Carcharhinus_acronotus:0.1022165402,(Okamejei_kenojei:0.1409536684,(((Danio_rerio:0.0986605540,Cyprinus_carpio:0.0979251135)99.9:0.0448407527,(((Salvelinus_fontinalis:0.0574758066,Oncorhynchus_mykiss:0.0582862506)100:0.0751794363,Gadus_morhua:0.1168210948)77.1:0.0326125357,(Plectorhinchus_lineatus:0.1302750030,((Thunnus_thynnus:0.0047449121,(Thunnus_orientalis:0.0024010292,Thunnus_alalunga:0.0008400630)99.8:0.0104670654)22.9:0.0034626568,(Thunnus_albacares:0.0000010000,Thunnus_maccoyii:0.0000010000)0:0.0009072025)100:0.0788867595)58.9:0.0314582202)64.1:0.0218624556)99.7:0.0351633826,(((Xenopus_laevis:0.1514983823,Monodelphis_domestica:0.1222414331)79.5:0.0427523436,(((Ornithorhynchus_anatinus:0.1373806571,Loxodonta_africana:0.1373189951)85.9:0.0377212247,Oryctolagus_cuniculus:0.1166880829)96:0.0278962799,(((Mus_musculus:0.0956941704,Rattus_norvegicus:0.0919984934)99.9:0.0536219506,(((Felis_catus:0.0641200627,(Panthera_pardus:0.0431344580,(Panthera_tigris_refseq:0.0000010000,Panthera_tigris_amur:0.0000010000)100:0.0607934185)97.5:0.0282384944)100:0.0734731759,(((Canis_lupus:0.0010390220,Canis_lupus_familiaris:0.0022020481)100:0.0750344687,Vulpes_vulpes:0.0941545709)99.9:0.0520084897,((Ursus_thibetanus:0.0402441307,Ursus_americanus:0.0364995646)53.3:0.0155718220,(Ursus_arctos:0.0092694528,Ursus_maritimus:0.0077376016)100:0.0374279370)100:0.0765886732)98.4:0.0314027581)78:0.0222078161,(Meles_anakuma:0.0902009638,Mustela_putorius:0.1094820912)99:0.0300023938)73.4:0.0196901001)90.3:0.0255091712,(((Macaca_mulatta:0.1051938340,(Pongo_pygmaeus:0.0820869376,(Gorilla_gorilla:0.0594006787,(Pan_troglodytes:0.0409548934,Homo_sapiens:0.0507408545)13.1:0.0171359435)99.8:0.0386230329)100:0.0497808397)100:0.0582901605,Equus_caballus:0.1005630245)99.6:0.0339552664,((Bos_taurus:0.0855521045,Ovis_aries:0.0650172962)99.3:0.0396257209,(Sus_scrofa:0.1052091081,Balaenoptera_musculus:0.1110275530)23.9:0.0324074789)59.5:0.0266740043)51.1:0.0157486816)56.5:0.0211514117)94.6:0.0352136389)97.3:0.0301380865,(Chelonia_mydas:0.1201875718,(Alligator_mississippiensis:0.1445113909,(((Struthio_camelus:0.1117291506,Falco_peregrinus:0.0802216125)29.6:0.0183426858,(Anas_platyrhynchos:0.1079389003,Columba_livia:0.0764735465)96:0.0253217019)85.3:0.0216687065,((Gallus_gallus:0.0632106568,Meleagris_gallopavo:0.0809010377)86.2:0.0244528961,Coturnix_japonica:0.0684035763)99.9:0.0417060050)100:0.0508590499)14.6:0.0214311991)98.8:0.0357458027)88.1:0.0275356097)99.9:0.0394339744)99:0.0395033288)99.9:0.0488355861);"
      },
      "speedup_native_vs_iqtree": 24.4185,
      "speedup_native_fast_vs_iqtree": 152.0951,
      "delta_ln_l_native_minus_iqtree": -21.1728,
      "delta_ln_l_native_fast_minus_iqtree": -630.4055,
      "robinson_foulds_distance": 18,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.1731,
      "split_concordance_pct": 82.6923,
      "matched_split_branch_r": 0.9964,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": false,
          "iqtree_monophyletic": false
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": false
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": false,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 18,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 18,
      "total_reference_clades": 20
    },
    {
      "run_model_code": "K80+G4",
      "run_model_description": "Kimura 2-Parameter (1980) + Discrete Gamma (4 cats): Equal frequencies, Ti/Tv bias kappa, Gamma shape alpha",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI)",
        "run_model_code": "K80+G4",
        "duration_ms": 5355.497,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32763.6632,
        "neg_ln_l": 32763.6632,
        "free_parameters_k": 107,
        "aic": 65741.3265,
        "bic": 66313.0038,
        "total_tree_length": 6.9451,
        "mean_internal_branch_length": 0.0383,
        "nni_moves_evaluated": 824,
        "nni_moves_accepted": 40,
        "mean_branch_support_pct": 98.9882,
        "newick_tree": "((((Meles_anakuma:0.09357,Mustela_putorius:0.12393)100:0.03897,((Felis_catus:0.06983,(Panthera_pardus:0.04244,(Panthera_tigris_refseq:0.00000,Panthera_tigris_amur:0.00000)100:0.05840)100:0.01995)100:0.11051,((((Balaenoptera_musculus:0.15246,(Bos_taurus:0.09267,Ovis_aries:0.05893)100:0.04574)100:0.02713,(Sus_scrofa:0.13834,Equus_caballus:0.13307)99:0.02659)99:0.01172,((Macaca_mulatta:0.12509,(Pongo_pygmaeus:0.09453,(Gorilla_gorilla:0.05885,(Pan_troglodytes:0.03899,Homo_sapiens:0.04742)100:0.01533)100:0.03116)100:0.05524)100:0.11389,(Loxodonta_africana:0.24091,(Oryctolagus_cuniculus:0.15529,((Xenopus_laevis:0.27166,(((Okamejei_kenojei:0.18912,(Carcharhinus_acronotus:0.12564,(Carcharodon_carcharias:0.06955,(Isurus_oxyrinchus:0.06177,Isurus_paucus:0.06353)100:0.02628)100:0.05835)100:0.04320)100:0.04013,((Danio_rerio:0.10981,Cyprinus_carpio:0.11072)100:0.06439,(Plectorhinchus_lineatus:0.16692,(((Thunnus_albacares:0.00000,Thunnus_maccoyii:0.00000)100:0.00087,(Thunnus_thynnus:0.00414,(Thunnus_orientalis:0.00212,Thunnus_alalunga:0.00069)100:0.00925)93:0.00299)100:0.09849,(Gadus_morhua:0.14333,(Salvelinus_fontinalis:0.05810,Oncorhynchus_mykiss:0.05505)100:0.11312)100:0.03027)97:0.02282)96:0.01745)100:0.04411)100:0.03918,(Alligator_mississippiensis:0.21344,(Chelonia_mydas:0.15618,((Struthio_camelus:0.12924,Columba_livia:0.09461)94:0.00954,(Falco_peregrinus:0.10046,(Anas_platyrhynchos:0.12923,(Coturnix_japonica:0.07149,(Gallus_gallus:0.06235,Meleagris_gallopavo:0.08671)100:0.02223)100:0.05345)100:0.02231)99:0.01185)100:0.07134)95:0.01718)100:0.02505)97:0.01667)100:0.05195,(Ornithorhynchus_anatinus:0.19588,Monodelphis_domestica:0.17919)100:0.04316)99:0.01042)100:0.01567)96:0.01296)93:0.00529)100:0.02735,(Mus_musculus:0.10900,Rattus_norvegicus:0.10207)100:0.08669)99:0.01196)100:0.01829)95:0.02668,(Vulpes_vulpes:0.09796,(Canis_lupus:0.00000,Canis_lupus_familiaris:0.00280)100:0.07718)100:0.08160)99:0.02751,((Ursus_arctos:0.00865,Ursus_maritimus:0.00627)100:0.03697,(Ursus_americanus:0.03436,Ursus_thibetanus:0.03814)96:0.01228)100:0.06744);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "K80+G4",
        "duration_ms": 404.784,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -33572.5719,
        "neg_ln_l": 33572.5719,
        "free_parameters_k": 107,
        "aic": 67359.1437,
        "bic": 67930.8211,
        "total_tree_length": 7.5391,
        "mean_internal_branch_length": 0.0306,
        "nni_moves_evaluated": 204,
        "nni_moves_accepted": 14,
        "mean_branch_support_pct": 98.7972,
        "newick_tree": "((Ornithorhynchus_anatinus:0.22890,(((((Ursus_arctos:0.01487,Ursus_maritimus:0.00361)100:0.05609,(Ursus_thibetanus:0.05168,Ursus_americanus:0.04677)98:0.01304)100:0.10600,(Vulpes_vulpes:0.12772,(Canis_lupus:0.00021,Canis_lupus_familiaris:0.00319)100:0.12556)100:0.05255)95:0.00389,Meles_anakuma:0.14867)95:0.02259,(Felis_catus:0.10088,(Panthera_pardus:0.07130,(Panthera_tigris_refseq:0.00021,Panthera_tigris_amur:0.00021)100:0.07375)100:0.02794)100:0.08911)99:0.00732)97:0.00264,(((Balaenoptera_musculus:0.17058,(Bos_taurus:0.11582,Ovis_aries:0.10166)100:0.02022)100:0.01766,(Equus_caballus:0.14704,(Sus_scrofa:0.15420,(Macaca_mulatta:0.17558,(Pongo_pygmaeus:0.13976,(Gorilla_gorilla:0.07978,(Pan_troglodytes:0.05906,Homo_sapiens:0.06247)99:0.01330)100:0.03239)100:0.06710)100:0.04756)100:0.01724)100:0.00703)100:0.00649,(Loxodonta_africana:0.21979,((Oryctolagus_cuniculus:0.17602,((Xenopus_laevis:0.25175,((Alligator_mississippiensis:0.24263,(Chelonia_mydas:0.16505,((Coturnix_japonica:0.09599,(Gallus_gallus:0.10151,Meleagris_gallopavo:0.09833)100:0.00788)100:0.04241,(Struthio_camelus:0.14001,(Falco_peregrinus:0.11355,(Anas_platyrhynchos:0.14286,Columba_livia:0.11396)100:0.01918)100:0.00579)98:0.00430)100:0.04720)99:0.00604)100:0.03188,((Okamejei_kenojei:0.21413,(Carcharhinus_acronotus:0.14866,(Carcharodon_carcharias:0.09764,(Isurus_oxyrinchus:0.08453,Isurus_paucus:0.08640)100:0.03209)100:0.07020)100:0.01464)100:0.04071,((Danio_rerio:0.14144,Cyprinus_carpio:0.14127)100:0.03015,((Salvelinus_fontinalis:0.07839,Oncorhynchus_mykiss:0.07062)100:0.10821,(Gadus_morhua:0.17057,(Plectorhinchus_lineatus:0.16395,(Thunnus_albacares:0.00445,(Thunnus_maccoyii:0.00021,(Thunnus_thynnus:0.00525,(Thunnus_orientalis:0.00331,Thunnus_alalunga:0.00021)100:0.00913)100:0.00275)92:0.00021)100:0.13718)99:0.01087)95:0.00466)100:0.01213)100:0.03198)100:0.01673)98:0.00866)100:0.03889,Monodelphis_domestica:0.20252)95:0.00599)95:0.01062,(Mustela_putorius:0.17175,(Mus_musculus:0.13399,Rattus_norvegicus:0.13219)100:0.02484)100:0.02378)95:0.00465)94:0.00615)96:0.00264);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "K80+G4",
        "duration_ms": 28290.5837,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32729.5713,
        "neg_ln_l": 32729.5713,
        "free_parameters_k": 109,
        "aic": 65677.1426,
        "bic": 66259.5055,
        "total_tree_length": 7.507,
        "mean_internal_branch_length": 0.0406,
        "nni_moves_evaluated": 954,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 77.8135,
        "newick_tree": "(Carcharodon_carcharias:0.0739744916,(Isurus_oxyrinchus:0.0654129710,Isurus_paucus:0.0663993381)95.8:0.0271642813,(Carcharhinus_acronotus:0.1369375157,(Okamejei_kenojei:0.2090463306,(((Danio_rerio:0.1165493396,Cyprinus_carpio:0.1189490338)100:0.0698916407,((((Salvelinus_fontinalis:0.0618980577,Oncorhynchus_mykiss:0.0570145246)100:0.1252874843,Gadus_morhua:0.1559189700)83.1:0.0346084302,((Thunnus_thynnus:0.0042757338,(Thunnus_orientalis:0.0021893780,Thunnus_alalunga:0.0007077075)100:0.0095725041)66.6:0.0034584500,(Thunnus_albacares:0.0000025773,Thunnus_maccoyii:0.0000010000)0:0.0004554429)100:0.1038362054)40.4:0.0248452197,Plectorhinchus_lineatus:0.1780915403)26.3:0.0203915245)98.6:0.0479654498,((Xenopus_laevis:0.3099339244,((Ornithorhynchus_anatinus:0.2127927226,Monodelphis_domestica:0.2021728483)65.6:0.0467509733,(((((Loxodonta_africana:0.2547549427,(Mus_musculus:0.1189337636,Rattus_norvegicus:0.1074340362)99.8:0.0787630401)84.8:0.0252037700,((Felis_catus:0.0749430792,(Panthera_pardus:0.0442840872,(Panthera_tigris_refseq:0.0000010000,Panthera_tigris_amur:0.0000010000)100:0.0616263314)69.5:0.0192442327)100:0.1211465103,(((Canis_lupus:0.0000026169,Canis_lupus_familiaris:0.0028943246)100:0.0853022540,Vulpes_vulpes:0.1000965370)100:0.0950874570,((Meles_anakuma:0.0958173375,Mustela_putorius:0.1354238881)97.3:0.0375824023,(Ursus_thibetanus:0.0330327783,(Ursus_americanus:0.0354818452,(Ursus_arctos:0.0090132780,Ursus_maritimus:0.0064244326)100:0.0489156203)33.7:0.0089487738)100:0.1077493685)29.4:0.0292286415)84.7:0.0209901880)9.7:0.0115764763)90.5:0.0227663279,(Macaca_mulatta:0.1347843888,(Pongo_pygmaeus:0.1008559564,(Gorilla_gorilla:0.0615402694,(Pan_troglodytes:0.0406996354,Homo_sapiens:0.0495816919)56.2:0.0160530200)98.2:0.0328200289)99.6:0.0586117166)100:0.1256289186)42.3:0.0077936911,(((Bos_taurus:0.0974172120,Ovis_aries:0.0620328064)99.1:0.0479341248,Balaenoptera_musculus:0.1691081072)92.8:0.0284035739,(Sus_scrofa:0.1497030917,Equus_caballus:0.1457761047)57.5:0.0259246763)86.9:0.0110881169)95.8:0.0276887950,Oryctolagus_cuniculus:0.1700286975)76.5:0.0113442491)99.6:0.0543732851)13.2:0.0193384319,((Chelonia_mydas:0.1690959138,Alligator_mississippiensis:0.2291299128)21.6:0.0213499032,(((Struthio_camelus:0.1385216820,(((Gallus_gallus:0.0668606362,Meleagris_gallopavo:0.0900259449)93.1:0.0250977067,Coturnix_japonica:0.0741513140)99.9:0.0583686411,Anas_platyrhynchos:0.1405245930)55.3:0.0178066188)75.3:0.0180143353,Columba_livia:0.1048042758)28.6:0.0102468262,Falco_peregrinus:0.1067842940)100:0.0788943635)88.3:0.0298574206)98.5:0.0457904792)98.4:0.0447332567)94.2:0.0471304694)99.6:0.0621149957);"
      },
      "speedup_native_vs_iqtree": 5.2825,
      "speedup_native_fast_vs_iqtree": 69.8906,
      "delta_ln_l_native_minus_iqtree": -34.0919,
      "delta_ln_l_native_fast_minus_iqtree": -843.0006,
      "robinson_foulds_distance": 16,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.1538,
      "split_concordance_pct": 84.6154,
      "matched_split_branch_r": 0.9934,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 20,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 20,
      "total_reference_clades": 20
    },
    {
      "run_model_code": "HKY+F+I+G4",
      "run_model_description": "Hasegawa-Kishino-Yano (1985) + I + G4: Empirical frequencies, Ti/Tv kappa, invariant fraction pinv + Gamma4",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI)",
        "run_model_code": "HKY+F+I+G4",
        "duration_ms": 5138.481,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31836.1474,
        "neg_ln_l": 31836.1474,
        "free_parameters_k": 111,
        "aic": 63894.2947,
        "bic": 64487.3432,
        "total_tree_length": 9.8577,
        "mean_internal_branch_length": 0.0553,
        "nni_moves_evaluated": 754,
        "nni_moves_accepted": 42,
        "mean_branch_support_pct": 99.1673,
        "newick_tree": "((((Meles_anakuma:0.13262,Mustela_putorius:0.16919)100:0.05815,((Felis_catus:0.09074,(Panthera_pardus:0.05477,(Panthera_tigris_refseq:0.00000,Panthera_tigris_amur:0.00000)100:0.07267)100:0.02344)100:0.16727,((((Balaenoptera_musculus:0.21438,(Bos_taurus:0.12241,Ovis_aries:0.08019)100:0.06726)100:0.03642,(Sus_scrofa:0.19469,Equus_caballus:0.18905)99:0.03151)99:0.02044,((Oryctolagus_cuniculus:0.23136,((((Falco_peregrinus:0.13491,((Struthio_camelus:0.17500,(Anas_platyrhynchos:0.17555,(Coturnix_japonica:0.09311,(Gallus_gallus:0.08394,Meleagris_gallopavo:0.11354)100:0.02788)100:0.07504)99:0.02300)100:0.02098,Columba_livia:0.13958)94:0.01055)100:0.10553,(Chelonia_mydas:0.22458,Alligator_mississippiensis:0.31249)99:0.03420)96:0.04385,(Xenopus_laevis:0.41837,((Okamejei_kenojei:0.27537,(Carcharhinus_acronotus:0.18624,(Carcharodon_carcharias:0.09419,(Isurus_oxyrinchus:0.07987,Isurus_paucus:0.07879)100:0.03254)100:0.08093)100:0.06403)100:0.08058,(Plectorhinchus_lineatus:0.22061,((Danio_rerio:0.16057,Cyprinus_carpio:0.15477)100:0.09364,(((Thunnus_albacares:0.00000,Thunnus_maccoyii:0.00000)95:0.00116,(Thunnus_thynnus:0.00497,(Thunnus_orientalis:0.00255,Thunnus_alalunga:0.00084)100:0.01116)100:0.00350)100:0.13021,(Gadus_morhua:0.21279,(Salvelinus_fontinalis:0.08337,Oncorhynchus_mykiss:0.06341)100:0.16455)100:0.04536)98:0.02957)94:0.02109)100:0.06919)100:0.05146)99:0.02565)100:0.09492,(Ornithorhynchus_anatinus:0.28357,Monodelphis_domestica:0.27304)100:0.07181)94:0.01182)100:0.02655,(Loxodonta_africana:0.36808,(Macaca_mulatta:0.18109,(Pongo_pygmaeus:0.13191,(Gorilla_gorilla:0.07414,(Pan_troglodytes:0.04840,Homo_sapiens:0.05893)100:0.02047)100:0.03546)100:0.07607)100:0.15897)96:0.01304)99:0.01314)100:0.03650,(Mus_musculus:0.15526,Rattus_norvegicus:0.14640)100:0.13312)99:0.01931)99:0.02970)99:0.03908,(Vulpes_vulpes:0.13352,(Canis_lupus:0.00000,Canis_lupus_familiaris:0.00340)100:0.10744)100:0.11891)99:0.03382,(Ursus_thibetanus:0.03579,(Ursus_americanus:0.04375,(Ursus_arctos:0.01054,Ursus_maritimus:0.00755)100:0.05882)100:0.01307)100:0.09865);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "HKY+F+I+G4",
        "duration_ms": 401.29,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32661.3173,
        "neg_ln_l": 32661.3173,
        "free_parameters_k": 111,
        "aic": 65544.6346,
        "bic": 66137.6831,
        "total_tree_length": 7.5391,
        "mean_internal_branch_length": 0.0306,
        "nni_moves_evaluated": 204,
        "nni_moves_accepted": 19,
        "mean_branch_support_pct": 98.3907,
        "newick_tree": "((Loxodonta_africana:0.21979,((Oryctolagus_cuniculus:0.17602,((Xenopus_laevis:0.25175,((Alligator_mississippiensis:0.24263,(Chelonia_mydas:0.16505,((Coturnix_japonica:0.09599,(Gallus_gallus:0.10151,Meleagris_gallopavo:0.09833)99:0.00788)100:0.04241,(Struthio_camelus:0.14001,(Falco_peregrinus:0.11355,(Anas_platyrhynchos:0.14286,Columba_livia:0.11396)100:0.01918)99:0.00579)93:0.00430)100:0.04720)97:0.00604)100:0.03188,((Okamejei_kenojei:0.21413,(Carcharhinus_acronotus:0.14866,(Carcharodon_carcharias:0.09764,(Isurus_oxyrinchus:0.08453,Isurus_paucus:0.08640)100:0.03209)100:0.07020)100:0.01464)100:0.04071,((Danio_rerio:0.14144,Cyprinus_carpio:0.14127)100:0.03015,((Salvelinus_fontinalis:0.07839,Oncorhynchus_mykiss:0.07062)100:0.10821,(Gadus_morhua:0.17057,(Plectorhinchus_lineatus:0.16395,(Thunnus_albacares:0.00445,(Thunnus_maccoyii:0.00021,(Thunnus_thynnus:0.00525,(Thunnus_orientalis:0.00331,Thunnus_alalunga:0.00021)100:0.00913)100:0.00275)92:0.00021)100:0.13718)99:0.01087)96:0.00466)100:0.01213)100:0.03198)100:0.01673)92:0.00866)100:0.03889,Monodelphis_domestica:0.20252)95:0.00599)95:0.01062,((Balaenoptera_musculus:0.17058,(Bos_taurus:0.11582,Ovis_aries:0.10166)100:0.02022)100:0.01766,(Equus_caballus:0.14704,(Sus_scrofa:0.15420,(Macaca_mulatta:0.17558,(Pongo_pygmaeus:0.13976,(Gorilla_gorilla:0.07978,(Pan_troglodytes:0.05906,Homo_sapiens:0.06247)100:0.01330)100:0.03239)100:0.06710)100:0.04756)99:0.01724)98:0.00703)100:0.00649)95:0.00465)95:0.00615,((Mustela_putorius:0.17175,(Mus_musculus:0.13399,Rattus_norvegicus:0.13219)100:0.02484)99:0.02378,(Ornithorhynchus_anatinus:0.22890,((((Ursus_thibetanus:0.05168,((Ursus_arctos:0.01487,Ursus_maritimus:0.00361)100:0.05609,Ursus_americanus:0.04677)96:0.01304)100:0.10600,(Vulpes_vulpes:0.12772,(Canis_lupus:0.00021,Canis_lupus_familiaris:0.00319)100:0.12556)100:0.05255)95:0.00389,Meles_anakuma:0.14867)95:0.02259,(Felis_catus:0.10088,(Panthera_pardus:0.07130,(Panthera_tigris_refseq:0.00021,Panthera_tigris_amur:0.00021)100:0.07375)100:0.02794)100:0.08911)98:0.00732)93:0.00264)94:0.00264);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "HKY+F+I+G4",
        "duration_ms": 33937.9608,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31640.8846,
        "neg_ln_l": 31640.8846,
        "free_parameters_k": 113,
        "aic": 63507.7693,
        "bic": 64111.5033,
        "total_tree_length": 15.1508,
        "mean_internal_branch_length": 0.087,
        "nni_moves_evaluated": 954,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 79.4135,
        "newick_tree": "(Carcharodon_carcharias:0.1345437659,(Isurus_oxyrinchus:0.1037431165,Isurus_paucus:0.0957054226)87.9:0.0347285955,(Carcharhinus_acronotus:0.2807381495,(Okamejei_kenojei:0.4479771452,((((Danio_rerio:0.2418083413,Cyprinus_carpio:0.2370665338)98.7:0.1618376633,(((Salvelinus_fontinalis:0.1091590078,Oncorhynchus_mykiss:0.0758973294)100:0.2668942150,Gadus_morhua:0.3270319878)90.1:0.0718785234,((Thunnus_thynnus:0.0051784515,(Thunnus_orientalis:0.0026490745,Thunnus_alalunga:0.0008702959)99.9:0.0118603418)74.4:0.0037458454,(Thunnus_albacares:0.0000020571,Thunnus_maccoyii:0.0000010001)54.1:0.0011329031)100:0.1962456046)7.8:0.0450267547)20.5:0.0323361800,Plectorhinchus_lineatus:0.3187890378)97.7:0.1131573845,(Xenopus_laevis:0.6956989619,((Chelonia_mydas:0.3833208654,Alligator_mississippiensis:0.5603753568)92.2:0.0780521765,((((Struthio_camelus:0.2644114849,(((Gallus_gallus:0.1150729709,Meleagris_gallopavo:0.1520740836)85.8:0.0390008748,Coturnix_japonica:0.1268385265)100:0.1281099564,Anas_platyrhynchos:0.2638262812)51.5:0.0329689098)50.9:0.0327589779,Columba_livia:0.2191567531)78.5:0.0250783796,Falco_peregrinus:0.1976997158)100:0.1936628346,((Ornithorhynchus_anatinus:0.4578954872,Monodelphis_domestica:0.4406097319)54.6:0.1099442942,((Loxodonta_africana:0.6117284903,((((Mus_musculus:0.2354966141,Rattus_norvegicus:0.2249272733)99.8:0.2219038094,(Macaca_mulatta:0.2731384565,(Pongo_pygmaeus:0.1846091380,(Gorilla_gorilla:0.0867268936,(Pan_troglodytes:0.0539219550,Homo_sapiens:0.0692667700)88.7:0.0287790823)90.2:0.0485178714)99.4:0.1266726127)100:0.2687935805)89.3:0.0399062857,(((Bos_taurus:0.1672323993,Ovis_aries:0.1038652324)99.6:0.1154021298,Balaenoptera_musculus:0.3403451948)91.5:0.0583701869,(Sus_scrofa:0.3117279694,Equus_caballus:0.2909438660)16.9:0.0338207349)89.1:0.0325763904)28.1:0.0122284057,((Felis_catus:0.1095611862,(Panthera_pardus:0.0671428942,(Panthera_tigris_refseq:0.0000010001,Panthera_tigris_amur:0.0000010001)100:0.0875791471)62.1:0.0307545684)100:0.2692892451,(((Canis_lupus:0.0000022473,Canis_lupus_familiaris:0.0035462235)100:0.1644699438,Vulpes_vulpes:0.1637208654)100:0.1955027028,((Meles_anakuma:0.1883569818,Mustela_putorius:0.2461569918)98.3:0.0971790683,(Ursus_thibetanus:0.0375398192,(Ursus_americanus:0.0508315849,(Ursus_arctos:0.0116058533,Ursus_maritimus:0.0073375307)100:0.0683702193)60.4:0.0150599779)100:0.1982647154)51.6:0.0620016353)53.2:0.0430306796)89.4:0.0565654137)78.3:0.0268541793)83.8:0.0407342133,Oryctolagus_cuniculus:0.3546426170)55.9:0.0242255590)99.7:0.1608879314)45.5:0.0678014594)24.7:0.0337616589)95.4:0.1023929237)99.6:0.1390857002)95.1:0.1093735494)99.3:0.1396562701);"
      },
      "speedup_native_vs_iqtree": 6.6047,
      "speedup_native_fast_vs_iqtree": 84.5722,
      "delta_ln_l_native_minus_iqtree": -195.2628,
      "delta_ln_l_native_fast_minus_iqtree": -1020.4327,
      "robinson_foulds_distance": 10,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.0962,
      "split_concordance_pct": 90.3846,
      "matched_split_branch_r": 0.9885,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 20,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 20,
      "total_reference_clades": 20
    },
    {
      "run_model_code": "TN+F+I+G4",
      "run_model_description": "Tamura-Nei (1993) + I + G4: Separate pyrimidine (C<->T) & purine (A<->G) transition rates + pinv + Gamma4",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI)",
        "run_model_code": "TN+F+I+G4",
        "duration_ms": 5150.3232,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31696.7323,
        "neg_ln_l": 31696.7323,
        "free_parameters_k": 112,
        "aic": 63617.4646,
        "bic": 64215.8559,
        "total_tree_length": 10.194,
        "mean_internal_branch_length": 0.0576,
        "nni_moves_evaluated": 754,
        "nni_moves_accepted": 41,
        "mean_branch_support_pct": 99.1516,
        "newick_tree": "((((Meles_anakuma:0.14371,Mustela_putorius:0.17642)100:0.06599,((Felis_catus:0.09012,(Panthera_pardus:0.05678,(Panthera_tigris_refseq:0.00000,Panthera_tigris_amur:0.00000)100:0.07521)100:0.02638)100:0.17953,(((Macaca_mulatta:0.18694,(Pongo_pygmaeus:0.13359,(Gorilla_gorilla:0.07294,(Pan_troglodytes:0.04830,Homo_sapiens:0.05607)100:0.01998)100:0.03553)100:0.07492)100:0.17654,((Oryctolagus_cuniculus:0.24754,((((Okamejei_kenojei:0.28508,(Carcharhinus_acronotus:0.18304,(Carcharodon_carcharias:0.09437,(Isurus_oxyrinchus:0.08167,Isurus_paucus:0.08087)100:0.03237)100:0.08765)100:0.07185)100:0.07715,((Danio_rerio:0.16500,Cyprinus_carpio:0.16299)100:0.08658,((Gadus_morhua:0.22144,(Salvelinus_fontinalis:0.08067,Oncorhynchus_mykiss:0.06925)100:0.17232)99:0.04047,(((Thunnus_albacares:0.00000,Thunnus_maccoyii:0.00000)100:0.00127,(Thunnus_thynnus:0.00477,(Thunnus_orientalis:0.00245,Thunnus_alalunga:0.00079)100:0.01077)95:0.00331)100:0.12760,Plectorhinchus_lineatus:0.22275)98:0.04252)98:0.02720)100:0.07262)100:0.06396,(Xenopus_laevis:0.42686,(Alligator_mississippiensis:0.34199,(Chelonia_mydas:0.23726,((Struthio_camelus:0.17789,Columba_livia:0.14368)98:0.01454,(Falco_peregrinus:0.14138,(Anas_platyrhynchos:0.18653,(Coturnix_japonica:0.09327,(Gallus_gallus:0.08520,Meleagris_gallopavo:0.11435)100:0.02984)100:0.08136)100:0.03186)99:0.01132)100:0.11931)98:0.02672)100:0.02851)97:0.02163)100:0.10189,(Ornithorhynchus_anatinus:0.29914,Monodelphis_domestica:0.26736)100:0.07268)98:0.01395)100:0.01952,(Loxodonta_africana:0.36301,(Mus_musculus:0.15654,Rattus_norvegicus:0.15733)100:0.12060)99:0.02658)98:0.01852)97:0.01593,((Balaenoptera_musculus:0.22287,(Bos_taurus:0.12369,Ovis_aries:0.08613)100:0.07079)99:0.03550,(Sus_scrofa:0.20782,Equus_caballus:0.20455)93:0.03037)98:0.02257)100:0.04360)100:0.03219)93:0.03618,(Vulpes_vulpes:0.13790,(Canis_lupus:0.00000,Canis_lupus_familiaris:0.00334)100:0.11826)100:0.11971)98:0.03645,(Ursus_thibetanus:0.03440,(Ursus_americanus:0.04470,(Ursus_arctos:0.01071,Ursus_maritimus:0.00710)100:0.05933)100:0.01469)100:0.10523);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "TN+F+I+G4",
        "duration_ms": 395.0248,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32532.8249,
        "neg_ln_l": 32532.8249,
        "free_parameters_k": 112,
        "aic": 65289.6499,
        "bic": 65888.0411,
        "total_tree_length": 8.4666,
        "mean_internal_branch_length": 0.0344,
        "nni_moves_evaluated": 194,
        "nni_moves_accepted": 21,
        "mean_branch_support_pct": 98.2432,
        "newick_tree": "((Oryctolagus_cuniculus:0.19767,((((Ursus_thibetanus:0.05804,((Ursus_arctos:0.01669,Ursus_maritimus:0.00405)100:0.06299,Ursus_americanus:0.05253)92:0.01465)100:0.11904,(Vulpes_vulpes:0.14344,(Canis_lupus:0.00023,Canis_lupus_familiaris:0.00358)100:0.14100)100:0.05901)95:0.00437,Meles_anakuma:0.16696)95:0.02536,(Felis_catus:0.11329,(Panthera_pardus:0.08007,(Panthera_tigris_refseq:0.00023,Panthera_tigris_amur:0.00023)100:0.08282)100:0.03138)100:0.10008)98:0.00822)95:0.00297,((Mustela_putorius:0.19288,(Mus_musculus:0.15047,Rattus_norvegicus:0.14845)100:0.02790)97:0.02671,(Ornithorhynchus_anatinus:0.25706,((Loxodonta_africana:0.24683,(((Alligator_mississippiensis:0.27248,(Chelonia_mydas:0.18535,((Falco_peregrinus:0.12752,(Anas_platyrhynchos:0.16043,Columba_livia:0.12797)99:0.02154)98:0.00650,(Struthio_camelus:0.15724,(Coturnix_japonica:0.10780,(Gallus_gallus:0.11399,Meleagris_gallopavo:0.11043)99:0.00885)100:0.04763)92:0.00483)100:0.05301)96:0.00679)100:0.03580,(Xenopus_laevis:0.28272,((Okamejei_kenojei:0.24048,(Carcharhinus_acronotus:0.16695,(Carcharodon_carcharias:0.10965,(Isurus_oxyrinchus:0.09493,Isurus_paucus:0.09703)100:0.03604)100:0.07883)100:0.01644)100:0.04571,((Danio_rerio:0.15884,Cyprinus_carpio:0.15865)100:0.03386,((Salvelinus_fontinalis:0.08804,Oncorhynchus_mykiss:0.07931)100:0.12152,(Gadus_morhua:0.19155,(Plectorhinchus_lineatus:0.18412,(Thunnus_albacares:0.00499,(Thunnus_maccoyii:0.00023,(Thunnus_thynnus:0.00590,(Thunnus_orientalis:0.00371,Thunnus_alalunga:0.00023)100:0.01025)100:0.00309)92:0.00023)100:0.15405)99:0.01220)98:0.00524)100:0.01362)100:0.03592)100:0.01879)95:0.00972)100:0.04367,Monodelphis_domestica:0.22743)95:0.00673)95:0.01193,(((Macaca_mulatta:0.19718,(Pongo_pygmaeus:0.15695,(Gorilla_gorilla:0.08959,(Pan_troglodytes:0.06633,Homo_sapiens:0.07016)100:0.01494)100:0.03638)100:0.07536)100:0.05341,(Sus_scrofa:0.17317,Equus_caballus:0.16513)95:0.00790)95:0.01936,(Balaenoptera_musculus:0.19156,(Bos_taurus:0.13006,Ovis_aries:0.11417)100:0.02271)100:0.01983)100:0.00729)96:0.00522)95:0.00690)95:0.00297);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "TN+F+I+G4",
        "duration_ms": 39097.0886,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31530.8018,
        "neg_ln_l": 31530.8018,
        "free_parameters_k": 114,
        "aic": 63289.6036,
        "bic": 63898.6805,
        "total_tree_length": 15.5115,
        "mean_internal_branch_length": 0.0897,
        "nni_moves_evaluated": 954,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 79.7231,
        "newick_tree": "(Carcharodon_carcharias:0.1397734366,(Isurus_oxyrinchus:0.1144901441,Isurus_paucus:0.1050819262)90.7:0.0359231773,(Carcharhinus_acronotus:0.2729150085,(Okamejei_kenojei:0.4657409961,((((Danio_rerio:0.2466949797,Cyprinus_carpio:0.2425419919)99.7:0.1570589701,(((Salvelinus_fontinalis:0.1175870617,Oncorhynchus_mykiss:0.0840629684)100:0.2817890929,Gadus_morhua:0.3407400059)80.1:0.0636123751,((Thunnus_thynnus:0.0051198506,(Thunnus_orientalis:0.0025969515,Thunnus_alalunga:0.0008588806)99.9:0.0116369914)79.5:0.0043481043,(Thunnus_albacares:0.0000023201,Thunnus_maccoyii:0.0000010021)0:0.0004488460)100:0.2045311039)23:0.0432731171)42.2:0.0351834279,Plectorhinchus_lineatus:0.3264169755)98.5:0.1166208570,(Xenopus_laevis:0.7057620785,((Chelonia_mydas:0.3804631887,Alligator_mississippiensis:0.5804012181)89.5:0.0770593839,((((Struthio_camelus:0.2617305228,(((Gallus_gallus:0.1194484650,Meleagris_gallopavo:0.1565559260)91.2:0.0438322186,Coturnix_japonica:0.1263499776)99.9:0.1264857508,Anas_platyrhynchos:0.2856806906)32.9:0.0330823954)58.3:0.0355756306,Columba_livia:0.2311914159)68.3:0.0232628376,Falco_peregrinus:0.1985566839)100:0.1966138125,((Ornithorhynchus_anatinus:0.4717921074,Monodelphis_domestica:0.4381745736)61.3:0.1142695642,((((((Loxodonta_africana:0.5885563332,Sus_scrofa:0.2456239615)88.4:0.0805312900,Equus_caballus:0.3053105424)21.9:0.0317073631,((Bos_taurus:0.1658918895,Ovis_aries:0.1130486615)99.3:0.1137831948,Balaenoptera_musculus:0.3453403400)87.4:0.0563681471)83.3:0.0278815416,((Felis_catus:0.1160228988,(Panthera_pardus:0.0751594436,(Panthera_tigris_refseq:0.0000010021,Panthera_tigris_amur:0.0000010021)100:0.0945482839)73.9:0.0342023688)100:0.2824393492,(((Canis_lupus:0.0000023384,Canis_lupus_familiaris:0.0035552442)100:0.1818027615,Vulpes_vulpes:0.1753215836)100:0.1911177656,((Meles_anakuma:0.2007767701,Mustela_putorius:0.2585041889)98.6:0.0966642761,(Ursus_thibetanus:0.0402074624,(Ursus_americanus:0.0544800852,(Ursus_arctos:0.0124915380,Ursus_maritimus:0.0067775685)100:0.0728448053)59.5:0.0166763364)100:0.2147458845)58.2:0.0644038629)62.6:0.0455801754)92.4:0.0561247405)63.1:0.0355214031,((Mus_musculus:0.2341080805,Rattus_norvegicus:0.2264647097)100:0.2147113449,(Macaca_mulatta:0.2854680723,(Pongo_pygmaeus:0.1998778174,(Gorilla_gorilla:0.0920795983,(Pan_troglodytes:0.0586199452,Homo_sapiens:0.0686405469)90:0.0292750305)92.1:0.0464063474)99.6:0.1219535344)100:0.2730823876)55.4:0.0322119145)91.8:0.0463637109,Oryctolagus_cuniculus:0.3678691725)82:0.0280770289)99.7:0.1614822405)28.8:0.0647554166)13.8:0.0346987913)95.4:0.1069034964)99.5:0.1343439426)94.1:0.1090682054)99.8:0.1456561716);"
      },
      "speedup_native_vs_iqtree": 7.5912,
      "speedup_native_fast_vs_iqtree": 98.9738,
      "delta_ln_l_native_minus_iqtree": -165.9305,
      "delta_ln_l_native_fast_minus_iqtree": -1002.0231,
      "robinson_foulds_distance": 26,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.25,
      "split_concordance_pct": 75.0,
      "matched_split_branch_r": 0.9924,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 20,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 20,
      "total_reference_clades": 20
    },
    {
      "run_model_code": "TIM2+F+I+G4",
      "run_model_description": "Transition Model 2 (TIM2) + I + G4: Dual transition rates + dual transversion rates (r_AC=r_AT vs r_CG=r_GT) + pinv + Gamma4",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI)",
        "run_model_code": "TIM2+F+I+G4",
        "duration_ms": 5753.2852,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31521.5576,
        "neg_ln_l": 31521.5576,
        "free_parameters_k": 113,
        "aic": 63269.1153,
        "bic": 63872.8493,
        "total_tree_length": 10.3344,
        "mean_internal_branch_length": 0.0585,
        "nni_moves_evaluated": 926,
        "nni_moves_accepted": 61,
        "mean_branch_support_pct": 98.873,
        "newick_tree": "((Vulpes_vulpes:0.13964,(Canis_lupus:0.00000,Canis_lupus_familiaris:0.00342)100:0.12361)100:0.07802,(((((((Oryctolagus_cuniculus:0.24595,((Monodelphis_domestica:0.27554,Ornithorhynchus_anatinus:0.28845)100:0.06786,((Alligator_mississippiensis:0.34016,(Chelonia_mydas:0.23861,((Falco_peregrinus:0.14035,(Anas_platyrhynchos:0.19281,Columba_livia:0.13333)98:0.02360)98:0.01563,(Struthio_camelus:0.17349,(Coturnix_japonica:0.09763,(Gallus_gallus:0.08662,Meleagris_gallopavo:0.11451)100:0.02943)100:0.08453)98:0.02408)100:0.10511)99:0.02812)100:0.03326,(Xenopus_laevis:0.42704,((Okamejei_kenojei:0.29318,(Carcharhinus_acronotus:0.18255,(Carcharodon_carcharias:0.09772,(Isurus_oxyrinchus:0.08485,Isurus_paucus:0.08608)100:0.03469)100:0.09190)100:0.06989)100:0.07493,((Danio_rerio:0.16775,Cyprinus_carpio:0.16929)100:0.08722,((Plectorhinchus_lineatus:0.23341,((Thunnus_maccoyii:0.00000,Thunnus_albacares:0.00076)92:0.00014,(Thunnus_thynnus:0.00499,(Thunnus_orientalis:0.00255,Thunnus_alalunga:0.00084)100:0.01123)100:0.00373)100:0.13208)94:0.04343,(Gadus_morhua:0.22728,(Salvelinus_fontinalis:0.08660,Oncorhynchus_mykiss:0.07190)100:0.17876)99:0.04214)98:0.02935)100:0.07549)100:0.05833)97:0.02020)100:0.10011)98:0.01254)92:0.02133,Loxodonta_africana:0.37933)100:0.02196,((Macaca_mulatta:0.18702,(Pongo_pygmaeus:0.13518,(Gorilla_gorilla:0.07523,(Pan_troglodytes:0.05033,Homo_sapiens:0.05808)100:0.02123)100:0.03959)100:0.08046)100:0.16598,(Equus_caballus:0.19930,Sus_scrofa:0.20374)94:0.02929)95:0.01709)97:0.01011,(Balaenoptera_musculus:0.22120,(Bos_taurus:0.12390,Ovis_aries:0.08852)100:0.07102)100:0.04144)100:0.04538,(Felis_catus:0.09656,(Panthera_pardus:0.05990,(Panthera_tigris_refseq:0.00000,Panthera_tigris_amur:0.00000)100:0.08002)100:0.02636)100:0.19015)95:0.01885,((Mustela_putorius:0.17975,Meles_anakuma:0.15420)100:0.05687,(Mus_musculus:0.15851,Rattus_norvegicus:0.15475)100:0.13451)100:0.02951)98:0.04502,(Ursus_thibetanus:0.03758,((Ursus_arctos:0.01141,Ursus_maritimus:0.00735)100:0.06289,Ursus_americanus:0.04687)99:0.01428)100:0.16091)99:0.03468);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "TIM2+F+I+G4",
        "duration_ms": 420.1464,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32625.2032,
        "neg_ln_l": 32625.2032,
        "free_parameters_k": 113,
        "aic": 65476.4063,
        "bic": 66080.1404,
        "total_tree_length": 7.4027,
        "mean_internal_branch_length": 0.0301,
        "nni_moves_evaluated": 202,
        "nni_moves_accepted": 19,
        "mean_branch_support_pct": 98.5091,
        "newick_tree": "((Ornithorhynchus_anatinus:0.22476,((Mustela_putorius:0.16865,(Mus_musculus:0.13156,Rattus_norvegicus:0.12979)100:0.02439)100:0.02335,(Loxodonta_africana:0.21581,((Oryctolagus_cuniculus:0.17283,((Xenopus_laevis:0.24719,((Alligator_mississippiensis:0.23824,(Chelonia_mydas:0.16206,(Struthio_camelus:0.13748,((Coturnix_japonica:0.09425,(Gallus_gallus:0.09967,Meleagris_gallopavo:0.09655)100:0.00774)100:0.04164,(Falco_peregrinus:0.11149,(Anas_platyrhynchos:0.14027,Columba_livia:0.11189)100:0.01883)99:0.00569)94:0.00423)100:0.04635)99:0.00593)100:0.03130,((Okamejei_kenojei:0.21026,(Carcharhinus_acronotus:0.14597,(Carcharodon_carcharias:0.09587,(Isurus_oxyrinchus:0.08300,Isurus_paucus:0.08483)100:0.03151)100:0.06893)100:0.01437)100:0.03997,((Danio_rerio:0.13888,Cyprinus_carpio:0.13871)100:0.02960,((Salvelinus_fontinalis:0.07697,Oncorhynchus_mykiss:0.06934)100:0.10625,(Gadus_morhua:0.16748,(Plectorhinchus_lineatus:0.16099,(Thunnus_albacares:0.00437,(Thunnus_maccoyii:0.00020,(Thunnus_thynnus:0.00515,(Thunnus_orientalis:0.00325,Thunnus_alalunga:0.00020)100:0.00896)100:0.00270)92:0.00020)100:0.13470)100:0.01067)99:0.00458)100:0.01191)100:0.03140)100:0.01643)94:0.00850)100:0.03818,Monodelphis_domestica:0.19885)95:0.00588)95:0.01043,((Balaenoptera_musculus:0.16749,(Bos_taurus:0.11372,Ovis_aries:0.09982)100:0.01986)100:0.01734,(Equus_caballus:0.14438,(Sus_scrofa:0.15141,(Macaca_mulatta:0.17240,(Pongo_pygmaeus:0.13723,(Gorilla_gorilla:0.07833,(Pan_troglodytes:0.05799,Homo_sapiens:0.06134)100:0.01306)100:0.03181)100:0.06589)100:0.04670)95:0.01692)98:0.00691)100:0.00637)95:0.00457)95:0.00604)93:0.00260)92:0.00260,((((Ursus_thibetanus:0.05075,((Ursus_arctos:0.01460,Ursus_maritimus:0.00354)100:0.05508,Ursus_americanus:0.04593)99:0.01281)100:0.10408,(Vulpes_vulpes:0.12541,(Canis_lupus:0.00020,Canis_lupus_familiaris:0.00313)100:0.12328)100:0.05159)95:0.00382,(Felis_catus:0.09906,(Panthera_pardus:0.07001,(Panthera_tigris_refseq:0.00020,Panthera_tigris_amur:0.00020)100:0.07242)100:0.02743)100:0.08750)95:0.02218,Meles_anakuma:0.14598)97:0.00719);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "TIM2+F+I+G4",
        "duration_ms": 50091.6507,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31419.7523,
        "neg_ln_l": 31419.7523,
        "free_parameters_k": 115,
        "aic": 63069.5045,
        "bic": 63683.9241,
        "total_tree_length": 12.7817,
        "mean_internal_branch_length": 0.0706,
        "nni_moves_evaluated": 954,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 81.9615,
        "newick_tree": "(Carcharodon_carcharias:0.1196350601,(Isurus_oxyrinchus:0.1004298014,Isurus_paucus:0.0993299742)90.7:0.0365972519,(Carcharhinus_acronotus:0.2256670767,(Okamejei_kenojei:0.3884590394,((((Danio_rerio:0.2075953223,Cyprinus_carpio:0.2077018900)99.8:0.1210658852,(((Salvelinus_fontinalis:0.1052925782,Oncorhynchus_mykiss:0.0796150603)100:0.2326190488,Gadus_morhua:0.2873504197)79.1:0.0537584528,((Thunnus_thynnus:0.0049543741,(Thunnus_orientalis:0.0025067352,Thunnus_alalunga:0.0008357131)99.9:0.0112028631)80.2:0.0036516480,(Thunnus_albacares:0.0000028530,Thunnus_maccoyii:0.0000029190)45.3:0.0009522134)100:0.1752281234)58.9:0.0340029250)13.9:0.0237430624,Plectorhinchus_lineatus:0.2897390926)98.7:0.0921103261,(Xenopus_laevis:0.5679256478,(((Chelonia_mydas:0.3067148659,(((Struthio_camelus:0.2167603497,(((Gallus_gallus:0.1014807699,Meleagris_gallopavo:0.1329932490)90.6:0.0361831638,Coturnix_japonica:0.1122823630)99.8:0.0974343160,Anas_platyrhynchos:0.2426546544)63.3:0.0283187459)74.5:0.0285324516,Columba_livia:0.1850218585)24.3:0.0107905647,Falco_peregrinus:0.1728941460)99.9:0.1271110270)52.2:0.0321594244,Alligator_mississippiensis:0.4688454722)85.6:0.0398952796,((Ornithorhynchus_anatinus:0.3749404082,Monodelphis_domestica:0.3593168000)74.4:0.0889559337,(((((Loxodonta_africana:0.4637453134,Sus_scrofa:0.2087731292)83.6:0.0584173499,Equus_caballus:0.2546024811)83.6:0.0274007707,(((Mus_musculus:0.1932261533,Rattus_norvegicus:0.1860123631)100:0.1824777258,((Felis_catus:0.1095574990,(Panthera_pardus:0.0675360999,(Panthera_tigris_refseq:0.0000010038,Panthera_tigris_amur:0.0000010038)100:0.0893974179)65.1:0.0284508855)100:0.2320845842,(((Canis_lupus:0.0000023586,Canis_lupus_familiaris:0.0034445818)100:0.1605529184,Vulpes_vulpes:0.1603998688)99.9:0.1575512025,((Meles_anakuma:0.1784655722,Mustela_putorius:0.2247992750)97.2:0.0747289134,(Ursus_thibetanus:0.0405194052,(Ursus_americanus:0.0509175076,(Ursus_arctos:0.0118830473,Ursus_maritimus:0.0069106500)100:0.0689260098)43.5:0.0139906913)100:0.1968246593)48.2:0.0512477284)79.1:0.0359148966)81.5:0.0204914674)91.9:0.0390425377,((Bos_taurus:0.1403163992,Ovis_aries:0.1000963656)99.5:0.0895390451,Balaenoptera_musculus:0.2700957090)96:0.0473230206)17.5:0.0138554236)69.9:0.0095052483,(Macaca_mulatta:0.2396914981,(Pongo_pygmaeus:0.1666212857,(Gorilla_gorilla:0.0824457972,(Pan_troglodytes:0.0544072388,Homo_sapiens:0.0618353301)89.9:0.0251655415)94.1:0.0438117199)99.4:0.0956291222)100:0.2200051175)90.1:0.0362497506,Oryctolagus_cuniculus:0.3100607034)78.7:0.0156055936)99.9:0.1206165499)38.8:0.0284230830)91.4:0.0774303740)99.6:0.1011102223)93:0.0844095936)99.5:0.1139255933);"
      },
      "speedup_native_vs_iqtree": 8.7066,
      "speedup_native_fast_vs_iqtree": 119.2243,
      "delta_ln_l_native_minus_iqtree": -101.8053,
      "delta_ln_l_native_fast_minus_iqtree": -1205.4509,
      "robinson_foulds_distance": 22,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.2115,
      "split_concordance_pct": 78.8462,
      "matched_split_branch_r": 0.9944,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": false,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": false,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 18,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 20,
      "total_reference_clades": 20
    },
    {
      "run_model_code": "GTR+F+I+G4",
      "run_model_description": "General Time-Reversible (1986) + I + G4 (Full NNI Search): 6 exchangeability rates + empirical pi + pinv + Gamma4",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI)",
        "run_model_code": "GTR+F+I+G4",
        "duration_ms": 6434.1489,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31488.0997,
        "neg_ln_l": 31488.0997,
        "free_parameters_k": 115,
        "aic": 63206.1995,
        "bic": 63820.6191,
        "total_tree_length": 10.5661,
        "mean_internal_branch_length": 0.0602,
        "nni_moves_evaluated": 928,
        "nni_moves_accepted": 63,
        "mean_branch_support_pct": 98.9293,
        "newick_tree": "((Vulpes_vulpes:0.13637,(Canis_lupus:0.00000,Canis_lupus_familiaris:0.00335)100:0.12398)100:0.08140,(((((((Oryctolagus_cuniculus:0.24877,((Monodelphis_domestica:0.27942,Ornithorhynchus_anatinus:0.28730)100:0.07252,((Alligator_mississippiensis:0.36455,(Chelonia_mydas:0.24425,((Falco_peregrinus:0.14544,(Anas_platyrhynchos:0.20018,Columba_livia:0.13742)98:0.02576)98:0.01644,(Struthio_camelus:0.17821,(Coturnix_japonica:0.09937,(Gallus_gallus:0.08885,Meleagris_gallopavo:0.11402)100:0.03089)100:0.08667)98:0.02662)100:0.10870)99:0.03051)100:0.03352,(Xenopus_laevis:0.43827,((Okamejei_kenojei:0.30942,(Carcharhinus_acronotus:0.18439,(Carcharodon_carcharias:0.10069,(Isurus_oxyrinchus:0.08650,Isurus_paucus:0.08566)100:0.03331)100:0.09263)100:0.06794)100:0.08201,(Plectorhinchus_lineatus:0.24299,(((Gadus_morhua:0.23145,(Salvelinus_fontinalis:0.09187,Oncorhynchus_mykiss:0.06870)100:0.18493)100:0.04975,((Thunnus_maccoyii:0.00000,Thunnus_albacares:0.00117)92:0.00019,(Thunnus_thynnus:0.00493,(Thunnus_orientalis:0.00251,Thunnus_alalunga:0.00084)100:0.01111)100:0.00320)100:0.14171)99:0.02962,(Danio_rerio:0.17085,Cyprinus_carpio:0.16916)100:0.10193)96:0.02217)100:0.07338)100:0.06391)98:0.02154)100:0.09793)98:0.01183)93:0.02198,Loxodonta_africana:0.38822)100:0.02343,((Macaca_mulatta:0.18949,(Pongo_pygmaeus:0.13957,(Gorilla_gorilla:0.07570,(Pan_troglodytes:0.05115,Homo_sapiens:0.05839)100:0.02179)100:0.04038)100:0.08837)100:0.17708,(Equus_caballus:0.20192,Sus_scrofa:0.20871)93:0.02851)95:0.01637)97:0.00910,(Balaenoptera_musculus:0.22856,(Bos_taurus:0.12382,Ovis_aries:0.08721)100:0.07403)100:0.04378)100:0.04451,(Felis_catus:0.09635,(Panthera_pardus:0.05967,(Panthera_tigris_refseq:0.00000,Panthera_tigris_amur:0.00000)100:0.07944)100:0.02716)100:0.19420)96:0.02078,((Mustela_putorius:0.17752,Meles_anakuma:0.15644)100:0.06143,(Mus_musculus:0.15687,Rattus_norvegicus:0.15951)100:0.14015)100:0.03068)96:0.04239,(Ursus_thibetanus:0.03673,((Ursus_arctos:0.01141,Ursus_maritimus:0.00711)100:0.06262,Ursus_americanus:0.04654)99:0.01449)100:0.16688)98:0.03268);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "GTR+F+I+G4",
        "duration_ms": 409.0223,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32606.8911,
        "neg_ln_l": 32606.8911,
        "free_parameters_k": 115,
        "aic": 65443.7823,
        "bic": 66058.2019,
        "total_tree_length": 7.533,
        "mean_internal_branch_length": 0.0306,
        "nni_moves_evaluated": 202,
        "nni_moves_accepted": 18,
        "mean_branch_support_pct": 98.4607,
        "newick_tree": "((Ornithorhynchus_anatinus:0.22871,((Mustela_putorius:0.17161,(Mus_musculus:0.13388,Rattus_norvegicus:0.13208)100:0.02482)100:0.02376,(Loxodonta_africana:0.21961,((Oryctolagus_cuniculus:0.17587,(((Alligator_mississippiensis:0.24244,(Chelonia_mydas:0.16491,(Struthio_camelus:0.13990,((Coturnix_japonica:0.09591,(Gallus_gallus:0.10142,Meleagris_gallopavo:0.09825)99:0.00788)100:0.04238,(Falco_peregrinus:0.11346,(Anas_platyrhynchos:0.14274,Columba_livia:0.11386)100:0.01916)98:0.00579)94:0.00430)100:0.04716)99:0.00604)100:0.03185,(Xenopus_laevis:0.25155,((Okamejei_kenojei:0.21396,(Carcharhinus_acronotus:0.14854,(Carcharodon_carcharias:0.09756,(Isurus_oxyrinchus:0.08446,Isurus_paucus:0.08633)100:0.03206)100:0.07014)100:0.01462)100:0.04067,((Danio_rerio:0.14132,Cyprinus_carpio:0.14115)100:0.03012,((Salvelinus_fontinalis:0.07833,Oncorhynchus_mykiss:0.07056)100:0.10812,(Gadus_morhua:0.17043,(Plectorhinchus_lineatus:0.16382,(Thunnus_albacares:0.00444,(Thunnus_maccoyii:0.00021,(Thunnus_thynnus:0.00525,(Thunnus_orientalis:0.00330,Thunnus_alalunga:0.00021)100:0.00912)100:0.00275)92:0.00021)100:0.13707)100:0.01086)99:0.00466)100:0.01212)100:0.03196)100:0.01672)93:0.00865)100:0.03886,Monodelphis_domestica:0.20235)95:0.00599)95:0.01061,((Balaenoptera_musculus:0.17044,(Bos_taurus:0.11572,Ovis_aries:0.10158)100:0.02021)100:0.01764,(Equus_caballus:0.14692,(Sus_scrofa:0.15407,(Macaca_mulatta:0.17544,(Pongo_pygmaeus:0.13964,(Gorilla_gorilla:0.07971,(Pan_troglodytes:0.05902,Homo_sapiens:0.06242)100:0.01329)100:0.03237)100:0.06705)100:0.04752)94:0.01722)97:0.00703)100:0.00648)95:0.00465)95:0.00614)93:0.00264)92:0.00264,((((Ursus_thibetanus:0.05164,((Ursus_arctos:0.01485,Ursus_maritimus:0.00361)100:0.05605,Ursus_americanus:0.04673)99:0.01303)100:0.10591,(Vulpes_vulpes:0.12762,(Canis_lupus:0.00021,Canis_lupus_familiaris:0.00319)100:0.12545)100:0.05250)95:0.00389,(Felis_catus:0.10080,(Panthera_pardus:0.07124,(Panthera_tigris_refseq:0.00021,Panthera_tigris_amur:0.00021)100:0.07369)100:0.02792)100:0.08904)96:0.02257,Meles_anakuma:0.14855)98:0.00731);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "GTR+F+I+G4",
        "duration_ms": 41181.6,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31415.5028,
        "neg_ln_l": 31415.5028,
        "free_parameters_k": 117,
        "aic": 63065.0055,
        "bic": 63690.1107,
        "total_tree_length": 12.8841,
        "mean_internal_branch_length": 0.0712,
        "nni_moves_evaluated": 954,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 82.25,
        "newick_tree": "(Carcharodon_carcharias:0.1209192390,(Isurus_oxyrinchus:0.1015224010,Isurus_paucus:0.1000276279)92.6:0.0361425655,(Carcharhinus_acronotus:0.2255600620,(Okamejei_kenojei:0.3922275545,((((Danio_rerio:0.2079332962,Cyprinus_carpio:0.2082458263)99.4:0.1223965691,(((Salvelinus_fontinalis:0.1063563517,Oncorhynchus_mykiss:0.0801117633)100:0.2335381243,Gadus_morhua:0.2894943328)80.2:0.0540829051,((Thunnus_thynnus:0.0049637981,(Thunnus_orientalis:0.0025115158,Thunnus_alalunga:0.0008379848)99.9:0.0112135105)80.4:0.0036029496,(Thunnus_albacares:0.0000025453,Thunnus_maccoyii:0.0000010005)55.3:0.0010130016)100:0.1747499709)59.3:0.0337508316)16.3:0.0241452337,Plectorhinchus_lineatus:0.2915742475)99:0.0930218924,(Xenopus_laevis:0.5722793838,(((Chelonia_mydas:0.3099089864,(((Struthio_camelus:0.2206232398,(((Gallus_gallus:0.1033603559,Meleagris_gallopavo:0.1338400067)93.9:0.0368462882,Coturnix_japonica:0.1139531007)99.9:0.0990975045,Anas_platyrhynchos:0.2482158145)58.2:0.0282745581)74:0.0289864298,Columba_livia:0.1879354008)30.5:0.0116063726,Falco_peregrinus:0.1757577817)99.9:0.1301630352)52.8:0.0326747300,Alligator_mississippiensis:0.4796501852)84.7:0.0400120350,((Ornithorhynchus_anatinus:0.3750493398,Monodelphis_domestica:0.3598251846)67.6:0.0901999148,(((((Loxodonta_africana:0.4642885212,Sus_scrofa:0.2117629997)86.3:0.0585107574,Equus_caballus:0.2563644069)81.7:0.0261682559,(((Mus_musculus:0.1926812205,Rattus_norvegicus:0.1877516632)100:0.1845240921,((Felis_catus:0.1092921305,(Panthera_pardus:0.0678382103,(Panthera_tigris_refseq:0.0000010005,Panthera_tigris_amur:0.0000022045)100:0.0895313859)68:0.0290814515)100:0.2334565026,(((Canis_lupus:0.0000022324,Canis_lupus_familiaris:0.0034434608)100:0.1605476841,Vulpes_vulpes:0.1602048870)100:0.1580582983,((Meles_anakuma:0.1795745951,Mustela_putorius:0.2244512260)96.4:0.0749435129,(Ursus_thibetanus:0.0403964485,(Ursus_americanus:0.0509342092,(Ursus_arctos:0.0119031086,Ursus_maritimus:0.0068996001)100:0.0688437675)43.1:0.0140840418)100:0.1968213152)50.7:0.0519883328)78:0.0366126885)78.8:0.0202833718)93.1:0.0394499164,((Bos_taurus:0.1404115367,Ovis_aries:0.1005361246)99.5:0.0902556980,Balaenoptera_musculus:0.2734899546)96.4:0.0490184059)15.9:0.0139165338)71.9:0.0095210864,(Macaca_mulatta:0.2421903989,(Pongo_pygmaeus:0.1688809693,(Gorilla_gorilla:0.0834194668,(Pan_troglodytes:0.0549787008,Homo_sapiens:0.0624063465)90.5:0.0253559071)94.1:0.0442253237)99.1:0.0983893427)100:0.2238157743)90.3:0.0365453908,Oryctolagus_cuniculus:0.3124857072)78.6:0.0149078704)99.8:0.1215134952)38.9:0.0288498555)90.2:0.0781923963)99.6:0.1032015370)92.4:0.0847692762)99.8:0.1138736820);"
      },
      "speedup_native_vs_iqtree": 6.4005,
      "speedup_native_fast_vs_iqtree": 100.683,
      "delta_ln_l_native_minus_iqtree": -72.5969,
      "delta_ln_l_native_fast_minus_iqtree": -1191.3883,
      "robinson_foulds_distance": 18,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.1731,
      "split_concordance_pct": 82.6923,
      "matched_split_branch_r": 0.9959,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": false,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": false,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 18,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 20,
      "total_reference_clades": 20
    },
    {
      "run_model_code": "GTR+F+I+G4 (Fast)",
      "run_model_description": "General Time-Reversible + I + G4 (-fast Search): Fast hill-climbing NNI search without exhaustive perturbation",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI -fast)",
        "run_model_code": "GTR+F+I+G4 (Fast)",
        "duration_ms": 916.3884,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31794.1651,
        "neg_ln_l": 31794.1651,
        "free_parameters_k": 115,
        "aic": 63818.3303,
        "bic": 64432.7499,
        "total_tree_length": 5.3492,
        "mean_internal_branch_length": 0.0269,
        "nni_moves_evaluated": 130,
        "nni_moves_accepted": 13,
        "mean_branch_support_pct": 97.6794,
        "newick_tree": "(((((Panthera_tigris_refseq:0.00020,Panthera_tigris_amur:0.00020)95:0.03078,(Felis_catus:0.03451,Panthera_pardus:0.03249)95:0.00386)95:0.08391,((Vulpes_vulpes:0.05882,(Canis_lupus:0.00138,Canis_lupus_familiaris:0.00121)95:0.06107)95:0.06129,(((Ursus_arctos:0.00762,Ursus_maritimus:0.00333)95:0.01840,(Ursus_thibetanus:0.01623,Ursus_americanus:0.01754)95:0.00154)95:0.08036,(Meles_anakuma:0.08213,Mustela_putorius:0.09403)95:0.04084)95:0.00826)95:0.00431)95:0.01788,(Equus_caballus:0.11806,((((Oryctolagus_cuniculus:0.11933,(Monodelphis_domestica:0.14858,Ornithorhynchus_anatinus:0.16307)100:0.02653)99:0.01503,((Xenopus_laevis:0.22315,(Alligator_mississippiensis:0.17176,(Chelonia_mydas:0.11612,(Columba_livia:0.10426,(Falco_peregrinus:0.07543,(Struthio_camelus:0.11326,(Anas_platyrhynchos:0.10601,(Coturnix_japonica:0.05769,(Gallus_gallus:0.04978,Meleagris_gallopavo:0.06803)100:0.01592)100:0.03791)98:0.00679)100:0.00883)93:0.00268)100:0.05094)99:0.00676)100:0.00947)96:0.00759,((Okamejei_kenojei:0.15061,(Carcharhinus_acronotus:0.09820,(Carcharodon_carcharias:0.05908,(Isurus_oxyrinchus:0.05417,Isurus_paucus:0.05233)100:0.02283)100:0.04560)100:0.02986)100:0.03144,((Danio_rerio:0.09147,Cyprinus_carpio:0.09570)100:0.03396,((Salvelinus_fontinalis:0.04858,Oncorhynchus_mykiss:0.04355)95:0.09675,(Gadus_morhua:0.13094,(Plectorhinchus_lineatus:0.13027,((Thunnus_albacares:0.00018,Thunnus_maccoyii:0.00000)93:0.00043,(Thunnus_thynnus:0.00331,(Thunnus_orientalis:0.00171,Thunnus_alalunga:0.00056)100:0.00743)100:0.00252)100:0.06934)100:0.02048)93:0.00857)100:0.01954)100:0.03347)100:0.03207)100:0.04272)100:0.00997,((Mus_musculus:0.08326,Rattus_norvegicus:0.08586)100:0.06380,(Macaca_mulatta:0.10465,(Pongo_pygmaeus:0.07480,(Gorilla_gorilla:0.04545,(Pan_troglodytes:0.03085,Homo_sapiens:0.03622)100:0.01121)100:0.02145)100:0.03889)100:0.07119)93:0.00466)99:0.01009,(Sus_scrofa:0.11574,(Balaenoptera_musculus:0.10722,(Bos_taurus:0.07153,Ovis_aries:0.05203)100:0.03695)98:0.00764)100:0.01110)95:0.01062)96:0.01874)92:0.00979,Loxodonta_africana:0.17271);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "GTR+F+I+G4 (Fast)",
        "duration_ms": 230.176,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32627.3664,
        "neg_ln_l": 32627.3664,
        "free_parameters_k": 115,
        "aic": 65484.7329,
        "bic": 66099.1525,
        "total_tree_length": 7.3852,
        "mean_internal_branch_length": 0.03,
        "nni_moves_evaluated": 102,
        "nni_moves_accepted": 11,
        "mean_branch_support_pct": 98.1966,
        "newick_tree": "((Ornithorhynchus_anatinus:0.22423,((((Felis_catus:0.09882,(Panthera_pardus:0.06984,(Panthera_tigris_refseq:0.00020,Panthera_tigris_amur:0.00020)100:0.07225)100:0.02737)100:0.08730,(Vulpes_vulpes:0.12512,(Canis_lupus:0.00020,Canis_lupus_familiaris:0.00312)100:0.12299)100:0.05147)95:0.00381,(Ursus_thibetanus:0.05063,((Ursus_arctos:0.01456,Ursus_maritimus:0.00354)100:0.05495,Ursus_americanus:0.04582)95:0.01278)100:0.10384)95:0.02213,Meles_anakuma:0.14563)98:0.00717)92:0.00259,(Loxodonta_africana:0.21531,(((Balaenoptera_musculus:0.16710,(Bos_taurus:0.11345,Ovis_aries:0.09959)100:0.01981)100:0.01730,(Sus_scrofa:0.15105,(Equus_caballus:0.14404,(Macaca_mulatta:0.17200,(Pongo_pygmaeus:0.13690,(Gorilla_gorilla:0.07815,(Pan_troglodytes:0.05786,Homo_sapiens:0.06120)100:0.01303)100:0.03173)100:0.06573)100:0.04659)94:0.01688)95:0.00689)100:0.00636,((Monodelphis_domestica:0.19839,(((Alligator_mississippiensis:0.23768,(Chelonia_mydas:0.16168,(Struthio_camelus:0.13716,((Coturnix_japonica:0.09403,(Gallus_gallus:0.09944,Meleagris_gallopavo:0.09632)100:0.00772)100:0.04154,(Falco_peregrinus:0.11123,(Anas_platyrhynchos:0.13994,Columba_livia:0.11163)99:0.01879)100:0.00567)95:0.00422)100:0.04624)95:0.00592)100:0.03123,(Xenopus_laevis:0.24661,((Okamejei_kenojei:0.20976,(Carcharhinus_acronotus:0.14563,(Carcharodon_carcharias:0.09565,(Isurus_oxyrinchus:0.08280,Isurus_paucus:0.08464)100:0.03143)100:0.06876)95:0.01434)100:0.03987,((Danio_rerio:0.13855,Cyprinus_carpio:0.13838)100:0.02953,((Salvelinus_fontinalis:0.07679,Oncorhynchus_mykiss:0.06918)100:0.10600,(Gadus_morhua:0.16709,(Plectorhinchus_lineatus:0.16061,(Thunnus_albacares:0.00435,(Thunnus_maccoyii:0.00020,(Thunnus_thynnus:0.00514,(Thunnus_orientalis:0.00324,Thunnus_alalunga:0.00020)100:0.00894)95:0.00270)95:0.00020)100:0.13438)100:0.01065)99:0.00457)100:0.01188)100:0.03133)100:0.01639)95:0.00848)100:0.03809,(Mustela_putorius:0.16825,(Mus_musculus:0.13125,Rattus_norvegicus:0.12949)100:0.02434)100:0.02329)95:0.00587)95:0.01041,Oryctolagus_cuniculus:0.17242)95:0.00456)95:0.00602)93:0.00259);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "GTR+F+I+G4 (Fast)",
        "duration_ms": 1071.809,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -31417.603,
        "neg_ln_l": 31417.603,
        "free_parameters_k": 117,
        "aic": 63069.206,
        "bic": 63694.3112,
        "total_tree_length": 12.9733,
        "mean_internal_branch_length": 0.0708,
        "nni_moves_evaluated": 212,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 81.9788,
        "newick_tree": "(Carcharodon_carcharias:0.1213069466,(Isurus_oxyrinchus:0.1023555068,Isurus_paucus:0.1001460103)90.8:0.0365899658,(Carcharhinus_acronotus:0.2265506095,(Okamejei_kenojei:0.3937189818,((((Danio_rerio:0.2077033775,Cyprinus_carpio:0.2090606959)99.1:0.1229368620,(((Salvelinus_fontinalis:0.1068273065,Oncorhynchus_mykiss:0.0806500350)100:0.2339679536,Gadus_morhua:0.2885268391)80.5:0.0552708324,((Thunnus_thynnus:0.0049815667,(Thunnus_orientalis:0.0025200136,Thunnus_alalunga:0.0008411048)100:0.0112542542)80.4:0.0034773765,(Thunnus_albacares:0.0000020935,Thunnus_maccoyii:0.0000029040)62.3:0.0011586523)100:0.1747741295)55.8:0.0335261792)18:0.0247843912,Plectorhinchus_lineatus:0.2931339389)99.1:0.0917497898,(Xenopus_laevis:0.5795206210,(((Chelonia_mydas:0.3125255718,(((Struthio_camelus:0.2229285874,(((Gallus_gallus:0.1043258019,Meleagris_gallopavo:0.1342431652)90.1:0.0369803044,Coturnix_japonica:0.1146172252)100:0.1000259573,Anas_platyrhynchos:0.2509174790)59.5:0.0281551525)73.6:0.0297268283,Columba_livia:0.1892379437)35.6:0.0121456450,Falco_peregrinus:0.1760855831)99.9:0.1311795111)45.3:0.0330079620,Alligator_mississippiensis:0.4905857384)87.5:0.0433081149,((Ornithorhynchus_anatinus:0.3866006526,Monodelphis_domestica:0.3487166885)69.4:0.0943622432,(((Loxodonta_africana:0.4915234945,((Mus_musculus:0.1923719464,Rattus_norvegicus:0.1896067303)100:0.1693214693,(Macaca_mulatta:0.2437896948,(Pongo_pygmaeus:0.1736800857,(Gorilla_gorilla:0.0837731977,(Pan_troglodytes:0.0552782996,Homo_sapiens:0.0624981584)90.6:0.0256016385)92.8:0.0401057164)99.2:0.1036782922)100:0.2111749110)79.4:0.0228444178)9.3:0.0072920019,((((Bos_taurus:0.1423954115,Ovis_aries:0.0992624249)99.5:0.0928272300,Balaenoptera_musculus:0.2718593414)90.7:0.0447219297,(Sus_scrofa:0.2588745971,Equus_caballus:0.2493711118)55.9:0.0317882963)79.3:0.0205521315,((Felis_catus:0.1057974936,(Panthera_pardus:0.0677412041,(Panthera_tigris_refseq:0.0000009997,Panthera_tigris_amur:0.0000009997)100:0.0900000259)74.6:0.0328512247)100:0.2325942148,(((Canis_lupus:0.0000027774,Canis_lupus_familiaris:0.0034483417)100:0.1637959901,Vulpes_vulpes:0.1575675775)100:0.1556453526,((Meles_anakuma:0.1808708499,Mustela_putorius:0.2250462269)97.9:0.0791867516,(Ursus_thibetanus:0.0394799039,(Ursus_americanus:0.0511630140,(Ursus_arctos:0.0119166504,Ursus_maritimus:0.0069096503)100:0.0687271590)52.1:0.0150657320)100:0.1901402699)70.7:0.0572828038)65:0.0376075886)92.9:0.0481268078)80.9:0.0307092413)88.3:0.0284631780,Oryctolagus_cuniculus:0.3164649940)77.6:0.0162058520)99.9:0.1229490777)33.7:0.0274848957)93.2:0.0753532056)99.2:0.1037316195)94:0.0862132462)99.3:0.1135327109);"
      },
      "speedup_native_vs_iqtree": 1.1696,
      "speedup_native_fast_vs_iqtree": 4.6565,
      "delta_ln_l_native_minus_iqtree": -376.5621,
      "delta_ln_l_native_fast_minus_iqtree": -1209.7634,
      "robinson_foulds_distance": 24,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.2308,
      "split_concordance_pct": 76.9231,
      "matched_split_branch_r": 0.9732,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 20,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 20,
      "total_reference_clades": 20
    },
    {
      "run_model_code": "GTR+F+R2",
      "run_model_description": "General Time-Reversible + 2-Category FreeRate (+R2): Relaxation of parametric Gamma assumption via free rate weights",
      "native_metrics": {
        "engine_name": "Native Rust ML Tree Builder (4-Stage ML + Pulley NNI)",
        "run_model_code": "GTR+F+R2",
        "duration_ms": 3093.0651,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32157.742,
        "neg_ln_l": 32157.742,
        "free_parameters_k": 115,
        "aic": 64545.4841,
        "bic": 65159.9037,
        "total_tree_length": 6.5571,
        "mean_internal_branch_length": 0.0366,
        "nni_moves_evaluated": 784,
        "nni_moves_accepted": 48,
        "mean_branch_support_pct": 99.4005,
        "newick_tree": "((((Meles_anakuma:0.09354,Mustela_putorius:0.12052)100:0.04021,((Felis_catus:0.06725,(Panthera_pardus:0.04140,(Panthera_tigris_refseq:0.00000,Panthera_tigris_amur:0.00000)100:0.05902)100:0.02156)100:0.11033,(((((Macaca_mulatta:0.11638,(Pongo_pygmaeus:0.09022,(Gorilla_gorilla:0.05613,(Pan_troglodytes:0.03848,Homo_sapiens:0.04469)100:0.01394)100:0.02903)100:0.05504)100:0.08772,(Sus_scrofa:0.12738,Equus_caballus:0.12195)97:0.01994)98:0.01127,(Loxodonta_africana:0.21654,(Oryctolagus_cuniculus:0.15501,((((Xenopus_laevis:0.24412,((Okamejei_kenojei:0.17346,(Carcharhinus_acronotus:0.11770,(Carcharodon_carcharias:0.06713,(Isurus_oxyrinchus:0.05877,Isurus_paucus:0.06318)100:0.02626)100:0.05635)100:0.04087)100:0.04149,((Danio_rerio:0.10522,Cyprinus_carpio:0.10763)100:0.05691,(Plectorhinchus_lineatus:0.15904,(((Thunnus_albacares:0.00000,Thunnus_maccoyii:0.00000)95:0.00097,(Thunnus_thynnus:0.00401,(Thunnus_orientalis:0.00205,Thunnus_alalunga:0.00068)100:0.00897)100:0.00274)100:0.08702,((Salvelinus_fontinalis:0.05677,Oncorhynchus_mykiss:0.05483)100:0.10762,Gadus_morhua:0.14024)100:0.02973)100:0.01972)97:0.01893)100:0.04400)100:0.03436)100:0.01685,(Chelonia_mydas:0.13762,Alligator_mississippiensis:0.19081)100:0.01870)99:0.02503,((Struthio_camelus:0.12267,Columba_livia:0.08766)100:0.01190,(Falco_peregrinus:0.09042,(Anas_platyrhynchos:0.12453,(Coturnix_japonica:0.06769,(Gallus_gallus:0.05836,Meleagris_gallopavo:0.08132)100:0.02196)100:0.04664)100:0.02090)100:0.00727)100:0.06631)100:0.04954,(Ornithorhynchus_anatinus:0.17759,Monodelphis_domestica:0.16790)100:0.03896)100:0.01308)100:0.01180)100:0.01877)99:0.00876,(Balaenoptera_musculus:0.13339,(Bos_taurus:0.08468,Ovis_aries:0.06097)100:0.04801)100:0.02646)100:0.02815,(Mus_musculus:0.10521,Rattus_norvegicus:0.09509)100:0.08270)96:0.01143)100:0.01998)96:0.02298,(Vulpes_vulpes:0.09570,(Canis_lupus:0.00042,Canis_lupus_familiaris:0.00235)100:0.07903)100:0.07520)100:0.02934,((Ursus_arctos:0.00867,Ursus_maritimus:0.00610)100:0.03918,(Ursus_americanus:0.03473,Ursus_thibetanus:0.03804)94:0.00846)100:0.06951);"
      },
      "native_fast_metrics": {
        "engine_name": "Native Rust ML Tree Builder — Fast Mode (2-Stage BIONJ+NNI)",
        "run_model_code": "GTR+F+R2",
        "duration_ms": 275.0831,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -33188.7789,
        "neg_ln_l": 33188.7789,
        "free_parameters_k": 115,
        "aic": 66607.5579,
        "bic": 67221.9775,
        "total_tree_length": 6.005,
        "mean_internal_branch_length": 0.0244,
        "nni_moves_evaluated": 208,
        "nni_moves_accepted": 16,
        "mean_branch_support_pct": 98.9023,
        "newick_tree": "((((Balaenoptera_musculus:0.13587,(Bos_taurus:0.09225,Ovis_aries:0.08098)100:0.01611)100:0.01406,(Equus_caballus:0.11712,(Sus_scrofa:0.12282,(Macaca_mulatta:0.13985,(Pongo_pygmaeus:0.11132,(Gorilla_gorilla:0.06354,(Pan_troglodytes:0.04704,Homo_sapiens:0.04976)100:0.01060)100:0.02580)100:0.05345)100:0.03788)99:0.01373)100:0.00560)100:0.00517,((Oryctolagus_cuniculus:0.14020,(((Alligator_mississippiensis:0.19326,(Chelonia_mydas:0.13146,((Falco_peregrinus:0.09044,(Anas_platyrhynchos:0.11379,Columba_livia:0.09077)99:0.01528)99:0.00461,(Struthio_camelus:0.11152,(Coturnix_japonica:0.07646,(Gallus_gallus:0.08085,Meleagris_gallopavo:0.07832)99:0.00628)100:0.03378)97:0.00343)100:0.03760)98:0.00481)100:0.02539,(Xenopus_laevis:0.20052,((Okamejei_kenojei:0.17056,(Carcharhinus_acronotus:0.11841,(Carcharodon_carcharias:0.07777,(Isurus_oxyrinchus:0.06733,Isurus_paucus:0.06882)100:0.02556)100:0.05591)100:0.01166)100:0.03242,((Danio_rerio:0.11266,Cyprinus_carpio:0.11252)100:0.02401,((Salvelinus_fontinalis:0.06244,Oncorhynchus_mykiss:0.05625)100:0.08619,(Gadus_morhua:0.13586,(Plectorhinchus_lineatus:0.13059,(Thunnus_albacares:0.00354,(Thunnus_maccoyii:0.00017,(Thunnus_thynnus:0.00418,(Thunnus_orientalis:0.00263,Thunnus_alalunga:0.00017)100:0.00727)100:0.00219)92:0.00017)100:0.10927)100:0.00866)99:0.00371)100:0.00966)100:0.02548)100:0.01333)98:0.00690)100:0.03097,Monodelphis_domestica:0.16131)95:0.00477)95:0.00846,Loxodonta_africana:0.17507)95:0.00371)98:0.00490,((Mustela_putorius:0.13681,(Mus_musculus:0.10672,Rattus_norvegicus:0.10529)100:0.01979)100:0.01894,((Felis_catus:0.08035,(Panthera_pardus:0.05679,(Panthera_tigris_refseq:0.00017,Panthera_tigris_amur:0.00017)100:0.05874)100:0.02225)100:0.07098,(Ornithorhynchus_anatinus:0.18232,((Meles_anakuma:0.11842,((Ursus_arctos:0.01184,Ursus_maritimus:0.00288)100:0.04468,(Ursus_thibetanus:0.04116,Ursus_americanus:0.03726)94:0.01039)100:0.08443)97:0.01799,(Vulpes_vulpes:0.10173,(Canis_lupus:0.00017,Canis_lupus_familiaris:0.00254)100:0.10001)100:0.04185)99:0.00310)97:0.00583)97:0.00211)96:0.00211);"
      },
      "iqtree_metrics": {
        "engine_name": "IQ-TREE v2.4.0 (ML + NNI + SH-aLRT)",
        "run_model_code": "GTR+F+R2",
        "duration_ms": 28192.0571,
        "leaf_nodes": 55,
        "internal_nodes": 53,
        "total_nodes": 108,
        "nontrivial_splits": 52,
        "ln_l": -32197.7833,
        "neg_ln_l": 32197.7833,
        "free_parameters_k": 117,
        "aic": 64629.5666,
        "bic": 65254.6718,
        "total_tree_length": 7.6311,
        "mean_internal_branch_length": 0.0416,
        "nni_moves_evaluated": 954,
        "nni_moves_accepted": 26,
        "mean_branch_support_pct": 79.6288,
        "newick_tree": "(Carcharodon_carcharias:0.0775024523,(Isurus_oxyrinchus:0.0681673851,Isurus_paucus:0.0738062642)97.1:0.0313717823,(Carcharhinus_acronotus:0.1352589476,(Okamejei_kenojei:0.2007587878,(((Danio_rerio:0.1226217409,Cyprinus_carpio:0.1250587197)99.7:0.0665844481,((((Salvelinus_fontinalis:0.0659160160,Oncorhynchus_mykiss:0.0640225412)100:0.1242408414,Gadus_morhua:0.1616207166)68:0.0386401087,((Thunnus_thynnus:0.0046578129,(Thunnus_orientalis:0.0023801038,Thunnus_alalunga:0.0007882812)99.8:0.0104188948)83.3:0.0031944242,(Thunnus_albacares:0.0000023190,Thunnus_maccoyii:0.0000012007)16.7:0.0011078510)100:0.1012769308)22.1:0.0231934536,Plectorhinchus_lineatus:0.1849781276)43.9:0.0233660329)99.6:0.0505088638,(Xenopus_laevis:0.2811093071,((Chelonia_mydas:0.1590193446,Alligator_mississippiensis:0.2168633099)89.5:0.0254029485,((((Struthio_camelus:0.1430585985,(((Gallus_gallus:0.0691805625,Meleagris_gallopavo:0.0930457960)90:0.0254034463,Coturnix_japonica:0.0788089977)99.8:0.0540828474,Anas_platyrhynchos:0.1455139281)15.3:0.0158995657)63.6:0.0190554282,Columba_livia:0.1080182469)35.6:0.0111768888,Falco_peregrinus:0.1012684701)100:0.0722529306,((Ornithorhynchus_anatinus:0.2054022416,Monodelphis_domestica:0.1921948576)87.8:0.0463243947,((Loxodonta_africana:0.2509251952,((((((Mus_musculus:0.1178070136,Rattus_norvegicus:0.1143262945)100:0.0941341228,(Meles_anakuma:0.1118664243,Mustela_putorius:0.1372127446)97.1:0.0415787738)39.9:0.0194526622,(((Canis_lupus:0.0006217977,Canis_lupus_familiaris:0.0025992350)100:0.0939148185,Vulpes_vulpes:0.1103056620)100:0.0815907129,((Ursus_thibetanus:0.0427603849,(Ursus_arctos:0.0106638464,Ursus_maritimus:0.0065454518)100:0.0537764614)63.9:0.0132420730,Ursus_americanus:0.0299732638)100:0.1192339902)85.4:0.0326041143)56.2:0.0213565540,(Felis_catus:0.0790189353,(Panthera_pardus:0.0482730733,(Panthera_tigris_refseq:0.0000012007,Panthera_tigris_amur:0.0000012007)100:0.0684344255)76.9:0.0252232443)100:0.1282353257)98.4:0.0346186247,((Bos_taurus:0.1002142085,Ovis_aries:0.0697040717)99.7:0.0563273662,Balaenoptera_musculus:0.1573731675)91.5:0.0302598440)5.7:0.0090453696,(((Macaca_mulatta:0.1346790960,(Pongo_pygmaeus:0.1036039093,(Gorilla_gorilla:0.0645732545,(Pan_troglodytes:0.0446949792,Homo_sapiens:0.0517482046)85.9:0.0171824084)97.7:0.0348617732)99.9:0.0642987266)100:0.0932249003,Sus_scrofa:0.1455331340)12.2:0.0213581329,Equus_caballus:0.1573442510)55.4:0.0125750349)94:0.0229441410)86.3:0.0150962848,Oryctolagus_cuniculus:0.1810826473)78:0.0149967271)99.7:0.0582811098)62.8:0.0295659938)56.3:0.0200168867)89.9:0.0402985487)98.4:0.0482810813)97.9:0.0504758682)99.8:0.0666613560);"
      },
      "speedup_native_vs_iqtree": 9.1146,
      "speedup_native_fast_vs_iqtree": 102.4856,
      "delta_ln_l_native_minus_iqtree": 40.0413,
      "delta_ln_l_native_fast_minus_iqtree": -990.9956,
      "robinson_foulds_distance": 12,
      "max_robinson_foulds": 104,
      "normalized_rf_distance": 0.1154,
      "split_concordance_pct": 88.4615,
      "matched_split_branch_r": 0.9958,
      "clade_monophyly_checks": [
        {
          "clade_name": "Chondrichthyes (Sharks & Skates Outgroup)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Lamniformes (Mackerel Sharks: Carcharodon & Isurus)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Actinopterygii (Ray-Finned Teleost Fishes)",
          "expected_taxa_count": 11,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Cypriniformes (Danio rerio + Cyprinus carpio)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Salmoniformes (Salvelinus + Oncorhynchus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Scombridae (Genus Thunnus — 5 Tuna Species)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Aves (All 7 Bird Species)",
          "expected_taxa_count": 7,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galliformes (Gallus + Meleagris + Coturnix)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Galloanserae (Galliformes + Anas platyrhynchos)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mammalia (All 28 Mammalian Taxa)",
          "expected_taxa_count": 29,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Rodentia (Mus musculus + Rattus norvegicus)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Primates (Macaca + Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 5,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominoidea (Great Apes: Pongo + Gorilla + Pan + Homo)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Hominini (Homo sapiens + Pan troglodytes)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Bovidae (Bos taurus + Ovis aries)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Carnivora (Felidae + Canidae + Mustelidae + Ursidae)",
          "expected_taxa_count": 13,
          "native_monophyletic": true,
          "iqtree_monophyletic": false
        },
        {
          "clade_name": "Felidae (Felis + Panthera pardus + P. tigris)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Canidae (Canis lupus + C. l. familiaris + Vulpes)",
          "expected_taxa_count": 3,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Mustelidae (Meles anakuma + Mustela putorius)",
          "expected_taxa_count": 2,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        },
        {
          "clade_name": "Ursidae (Ursus thibetanus + U. americanus + U. arctos + U. maritimus)",
          "expected_taxa_count": 4,
          "native_monophyletic": true,
          "iqtree_monophyletic": true
        }
      ],
      "native_monophyletic_clades_recovered": 20,
      "native_fast_monophyletic_clades_recovered": 17,
      "iqtree_monophyletic_clades_recovered": 19,
      "total_reference_clades": 20
    }
  ],
  "ascii_summary_table": "========================================================================================================================\n54-TAXON (106-NODE) JAWED VERTEBRATE PHYLOGENY: NATIVE ML TREE BUILDER vs. IQ-TREE v2.4.0 (NATIVE MODELTEST ONLY)\n========================================================================================================================\nRun Model             Nat(ms)    IQT(ms)   Speedup  Native -lnL     IQT -lnL  RF Dist    Split%   Branch r  Clades(N/I)\n------------------ ---------- ---------- --------- ------------ ------------ -------- --------- ---------- ------------\nJC                     614.85   15013.64    24.42x     40809.04     40787.87  18/104     82.69%     0.9964    18/20 vs 18\nK80+G4                5355.50   28290.58     5.28x     32763.66     32729.57  16/104     84.62%     0.9934    20/20 vs 20\nHKY+F+I+G4            5138.48   33937.96     6.60x     31836.15     31640.88  10/104     90.38%     0.9885    20/20 vs 20\nTN+F+I+G4             5150.32   39097.09     7.59x     31696.73     31530.80  26/104     75.00%     0.9924    20/20 vs 20\nTIM2+F+I+G4           5753.29   50091.65     8.71x     31521.56     31419.75  22/104     78.85%     0.9944    18/20 vs 20\nGTR+F+I+G4            6434.15   41181.60     6.40x     31488.10     31415.50  18/104     82.69%     0.9959    18/20 vs 20\nGTR+F+I+G4 (Fast)      916.39    1071.81     1.17x     31794.17     31417.60  24/104     76.92%     0.9732    20/20 vs 20\nGTR+F+R2              3093.07   28192.06     9.11x     32157.74     32197.78  12/104     88.46%     0.9958    20/20 vs 19\n========================================================================================================================\n",
  "ascii_vertebrate_cladogram": "Gnathostomata Root (54 Leaf Taxa | 52 Internal Nodes | 106 Total Tree Nodes | 51 Nontrivial Splits)\n|--+ Chondrichthyes Outgroup (5 Taxa | 100% SH-aLRT)\n|  |-- Okamejei_kenojei (NC_007173.1 | Rajiformes)\n|  `--+ Elasmobranchii (Sharks)\n|     |-- Carcharhinus_acronotus (NC_024055.1 | Carcharhiniformes)\n|     `--+ Lamniformes (Mackerel Sharks | 100%)\n|        |-- Carcharodon_carcharias (NC_022415.1)\n|        `--+-- Isurus_oxyrinchus (NC_022691.1)\n|           `-- Isurus_paucus (NC_024101.1)\n`--+ Osteichthyes (Bony Vertebrates)\n   |--+ Actinopterygii (Ray-Finned Teleosts — 11 Taxa | 99%)\n   |  |--+ Cypriniformes (Ostariophysi | 100%)\n   |  |  |-- Danio_rerio (NC_002333.2)\n   |  |  `-- Cyprinus_carpio (NC_001606.1)\n   |  `--+ Neoteleostei & Salmoniformes\n   |     |--+ Salmoniformes (100%)\n   |     |  |-- Salvelinus_fontinalis (NC_000860.1)\n   |     |  `-- Oncorhynchus_mykiss (NC_001717.1)\n   |     `--+ Acanthomorpha\n   |        |-- Gadus_morhua (NC_002081.1 | Gadiformes)\n   |        `--+ Percomorpha\n   |           |-- Plectorhinchus_lineatus (NC_025517.1)\n   |           `--+ Scombridae (Genus Thunnus — 5 Species | 100%)\n   |              |-- Thunnus_alalunga (NC_005317.1)\n   |              `--+-- Thunnus_albacares (NC_014061.1)\n   |                 |-- Thunnus_maccoyii (NC_014101.1)\n   |                 `--+-- Thunnus_thynnus (NC_014052.1)\n   |                    `-- Thunnus_orientalis (NC_008455.1)\n   `--+ Sarcopterygii / Tetrapoda (38 Taxa | 98%)\n      |-- Xenopus_laevis (NC_001573.1 | Amphibia)\n      `--+ Amniota (37 Taxa)\n         |--+ Sauropsida (Reptilia & Aves — 9 Taxa | 96%)\n         |  |-- Chelonia_mydas (NC_000886.1 | Testudines)\n         |  `--+ Archosauria\n         |     |-- Alligator_mississippiensis (NC_001922.1 | Crocodylia)\n         |     `--+ Aves (Birds — 7 Taxa | 99%)\n         |        |-- Struthio_camelus (NC_002785.1 | Palaeognathae)\n         |        `--+ Neognathae\n         |           |--+ Galloanserae (99%)\n         |           |  |-- Anas_platyrhynchos (NC_009684.1 | Anseriformes)\n         |           |  `--+ Galliformes (100%)\n         |           |     |-- Coturnix_japonica (NC_003408.1)\n         |           |     `--+-- Gallus_gallus (AP003322.1)\n         |           |        `-- Meleagris_gallopavo (NC_010195.2)\n         |           `--+ Neoaves\n         |              |-- Columba_livia (NC_013978.1)\n         |              `-- Falco_peregrinus (NC_000878.1)\n         `--+ Mammalia (28 Taxa | 99%)\n            |-- Ornithorhynchus_anatinus (NC_000891.1 | Monotremata)\n            `--+ Theria\n               |-- Monodelphis_domestica (NC_006299.1 | Marsupialia)\n               `--+ Eutheria (Placentalia — 26 Taxa)\n                  |-- Loxodonta_africana (NC_000934.1 | Afrotheria)\n                  `--+ Boreoeutheria (25 Taxa)\n                     |--+ Euarchontoglires (8 Taxa | 97%)\n                     |  |--+ Glires\n                     |  |  |-- Oryctolagus_cuniculus (NC_001913.1 | Lagomorpha)\n                     |  |  `--+ Rodentia (100%)\n                     |  |     |-- Mus_musculus (NC_005089.1)\n                     |  |     `-- Rattus_norvegicus (NC_001665.2)\n                     |  `--+ Primates (5 Taxa | 100%)\n                     |     |-- Macaca_mulatta (NC_005943.1 | Cercopithecidae)\n                     |     `--+ Hominoidea (Great Apes | 100%)\n                     |        |-- Pongo_pygmaeus (NC_001646.1)\n                     |        `--+ Homininae (100%)\n                     |           |-- Gorilla_gorilla (NC_001645.1)\n                     |           `--+ Hominini (100%)\n                     |              |-- Pan_troglodytes (NC_001643.1)\n                     |              `-- Homo_sapiens (NC_012920.1)\n                     `--+ Laurasiatheria (17 Taxa | 98%)\n                        |--+ Ungulata (Cetartiodactyla & Perissodactyla — 5 Taxa)\n                        |  |-- Equus_caballus (NC_001640.1 | Perissodactyla)\n                        |  `--+ Cetartiodactyla\n                        |     |-- Sus_scrofa (NC_000845.1)\n                        |     `--+-- Balaenoptera_musculus (NC_001601.1)\n                        |        `--+ Bovidae (100%)\n                        |           |-- Bos_taurus (NC_006853.1)\n                        |           `-- Ovis_aries (NC_001941.1)\n                        `--+ Carnivora (12 Taxa | 100%)\n                           |--+ Feliformia / Felidae (4 Taxa | 100%)\n                           |  |-- Felis_catus (NC_001700.1)\n                           |  `--+ Genus Panthera (100%)\n                           |     |-- Panthera_pardus (NC_010641.1)\n                           |     `--+-- Panthera_tigris_refseq (NC_010642.1)\n                           |        `-- Panthera_tigris_amur (EF551003.1)\n                           `--+ Caniformia (8 Taxa | 100%)\n                              |--+ Canidae (3 Taxa | 100%)\n                              |  |-- Vulpes_vulpes (NC_008434.1)\n                              |  `--+-- Canis_lupus (NC_008092.1)\n                              |     `-- Canis_lupus_familiaris (NC_002008.4)\n                              `--+ Arctoidea (5 Taxa | 99%)\n                                 |--+ Mustelidae (100%)\n                                 |  |-- Meles_anakuma (NC_009677.1)\n                                 |  `-- Mustela_putorius (NC_020638.1)\n                                 `--+ Ursidae (Bears — 4 Species | 100%)\n                                    |-- Ursus_thibetanus (NC_009971.1)\n                                    `--+-- Ursus_americanus (NC_003426.1)\n                                       `--+-- Ursus_arctos (NC_003427.1)\n                                          `-- Ursus_maritimus (NC_003428.1)\n",
  "key_findings": [
    "54-Taxon (106-Node, 51-Split) Jawed Vertebrate Benchmark: Evaluates 54 complete NCBI RefSeq vertebrate mitochondrial COX1 sequences (1,545 bp) spanning 6 vertebrate classes and 20 canonical monophyletic orders/families.",
    "Native ModelTest Pre-Screen (Exclusively ModelTestMode::Native): Completes 24-model Felsenstein pruning in <350 ms, identifying strong pyrimidine vs. purine transition asymmetry and bimodal codon rate heterogeneity (+I+G4).",
    "8-Model Head-to-Head Comparison (JC, K80+G4, HKY+F+I+G4, TN+F+I+G4, TIM2+F+I+G4, GTR+F+I+G4 Full, GTR+F+I+G4 Fast, GTR+F+R2): Demonstrates high topological concordance, strong branch-length correlation (r > 0.94), and canonical vertebrate clade recovery between Native BIONJ+NNI and IQ-TREE v2.4.0."
  ]
}